Revision of the Genus Pandanus Stickman, Part 17 
Species, Mostly New, in Borneo, Cambodia, and Vietnam 
Harold St. John 1 
In Borneo there are numerous known spe- 
cies of Pandanus , while in Cambodia there are 
but few. Several new ones from those regions 
are here proposed. Two new ones from Vietnam 
are also included, as are expanded descriptions 
and new locality records for certain species of 
Borneo and of Vietnam. 
SPECIES FROM BORNEO 
SECTION Aero stigma 
Pandanus pumilus sp. nov. (sect. Aero stigma). 
Fig. 206 
DIAGNOSIS HOLOTYPI: Licet acaulescens, foliis 
63-89 cm longis 7-8 mm latis subcoriaceis 
supra viridibus licet infra pallidioribus late 
sulcatis et cum plicis binis rotundatis inermibus, 
in sectione mediali cum 12-13 nervis secunda- 
riis parallelis in quoque latere, nervis tertiis 
nullis laminis gradatim in apice 15 cm longo 
trigono subulato diminuentibus eo 10 cm ex 
apice 3 mm lato, basi amplexicauli inermi 
purpurea excrassa et valde venosa, ex 4-4.5 cm 
marginibus cum aculeis 1—1.5 mm longis 1—5 
mm separatis subulatis compressis subadscen- 
dentibus pallidis, midnervo infra ultra mediam 
inermi, in sectione mediali marginibus cum 
serrulis 0 . 3 - 0. 5 mm longis 2-5 mm separatis 
adpressis, in apice subulato marginibus et mid- 
nervo infra cum serrulis 0.2-0. 3 mm longis 0.5— 
3 mm separatis, ad apicem supra plicis cum 
serrulis 0.2-0. 3 mm longis 4-5 mm separatis, 
inflorescentia foeminea erecta striata infra nuda 
parte I /3 infera retrorse scabra, bractea supera 
12 cm longo parte Va infera 8 mm lata lanceo- 
lata foliacea parte 44 supera trigona subulata, 
inflorescentia spicata cum 4-5 syncarpiis quaque 
1 B. P. Bishop Museum, Honolulu 17, Hawaii. 
Manuscript received October 17, 1961. 
sustenta cum bractea foliosa, ea mediali 2.3 cm 
longa 1 cm lata lanceolata, ultra mediam mar- 
ginibus et midnervo infra cum serrulis, syn- 
carpiis 1 . 5-2 cm longis 1 . 3 - 1. 5 cm diametro 
elliptico-subgloboso cum 113-160 drupis, eis 
6-7 mm longis 2. 5-3. 5 mm latis 1.5-3 mm 
crassis corpore oblanceo-ellipsoideo 5-6 mm 
longo stramineo valde striato, pileo cum basi 
1 mm alto depresse pyramidal! 5 - 6 -anguloso 
luteo et cum vicinis connatis, stylo 1 - 1.5 mm 
alto parte basali lanceoloidea apice in angulo 
recto crasse subulato, stigmate 1 - 1.5 mm longo 
distali late lineari nigro papillose paene in apice 
extento, endocarpio mediali ellipsoideo 3.8 mm 
longo cartilagineo stramineo lateribus 0.03 mm 
crassis, semine 3 mm longo ellipsoideo, meso- 
carpio apical i minuto, mesocarpio basali 1 mm 
longo in lateribus fibroso in media carnoso. 
DESCRIPTION OF ALL SPECIMENS EXAMINED: 
Apparently acaulescent, or perhaps with short 
slender stems; leaves 63-89 cm long, 7-8 mm 
wide, thin coriaceous, green above, apparently 
slightly paler beneath, broad-furrowed above 
the midrib and with two rounded pleats, above 
unarmed, at midsection with 12 or 13 parallel 
secondary veins in each half, no visible tertiary 
cross veins, very gradually narrowed to a trig- 
onous subulate apex 15 cm long, this 10 cm 
down 3 mm wide, the base amplexicaul, un- 
armed, purple, thin and strongly veiny, begin- 
ning 4-4.5 cm from the base the margins with 
prickles 1 - 1.5 mm long, 1-5 mm apart, flattened 
subulate, slightly ascending, pale; the midrib 
unarmed to beyond the middle; at midsection 
the margins with appressed serrulations 0.3-0. 5 
mm long, 2-5 mm apart; on the subulate apex 
the margins and midrib below with serrulations 
0 . 2 - 0. 3 mm long, 0 . 5-3 mm apart; towards the 
apex on the upper side the pleats with serrula- 
tions 0.2-0. 3 mm long, 4-5 mm apart; pis- 
tillate inflorescence erect, striate, naked except 
96 
