Page 334: Revision of Pandanus, 17. Borneo, Cambodia, Vietnam — St. John 
99 
John has the syncarp 20-26 cm long, 4.5 cm in 
diameter; drupes 9-10 mm long, 1.7-2. 5 mm 
wide; stigmas 0.9-1. 3 mm wide; and the leaves 
11 cm wide. 
The new epithet is the Latin adjective trig- 
onus, three-angled, and is given in reference to 
the three-angled syncarp. 
SECTION Pandanus 
Pandanus tectorius Soland. var. borneensis Mar- 
telli, Webbia 4(2) :4l0, pi 19, fig. 6, 
1914 
Fig. 208 
EXPANDED DESCRIPTION: From Wood & 
Wyatt-Smith A4,567: Shrub, 8.6 m tall; branch 
apex 3.5 cm in diameter; leaves 84-100 cm 
long, 4 cm wide, coriaceous, broad furrowed 
above the midrib, the sides arched and recurv- 
ing, at midsection with 46 parallel secondary 
veins in each half, but no visible tertiary veins, 
blade sword-shaped, tapering gradually to the 
trigonous, broad subulate apex, this 10 cm 
down 10 mm wide, the base amplexicaul, un- 
armed, but beginning 8 cm up the margins with 
prickles 2-3.5 mm long, 7-13 mm apart, arcuate 
subulate, ascending, pale; the midrib below be- 
ginning at 7-10 cm with prickles 3-3.5 mm 
long, 17-32 mm apart, heavy subulate, arcuate, 
reflexed; at midsection the margins with prickles 
2-4 mm long, 12-20 mm apart, similar, ascend- 
ing; the midrib below with prickles 2. 5-3.2 mm 
long, 35-39 mm apart, similar; on the subulate 
apex the margins with serrulations 0.2-0.3 mm 
long, 2-4 mm apart; the midrib below with 
similar serrulations but 3-6 mm apart; pha- 
langes 5. 3-5.8 cm long, 2.9-3. 5 cm wide, 2.5- 
2.9 cm thick, pyriform, 4-5 -angled, the sides 
gently convex, smooth, shining, when dried 
light brown, upper lA free, the apex convex, 
lateral sutures none, central apical sinuses 1-2 
mm deep, very shallow but the bottom half V- 
shaped; carpels 6 or 7, the apices low obtuse 
and the marginal ones with a small concavity 
distal of the stigma, the outer carpels 11/2-2 
times the larger; stigma 3-3.5 mm long, elliptic, 
creased, dark brown, elevated; proximal sinus 
a wide crack running V 5 —V 2 way to valley bot- 
tom; endocarp slightly supramedian, 22 mm 
long, bony, the inner parts dark mahogany 
colored, the outer parts pale, the lateral walls 
4-5 mm thick, the inner surfaces shining; seeds 
14-16 mm long, 3-4 mm in diameter, ellipsoid 
or obliquely so; upper mesocarp forming in each 
carpel apex a cavity with brownish medullary 
membranes; basal mesocarp fibrous and fleshy. 
SPECIMENS EXAMINED: North Borneo, Sipi- 
tang District, Sibubu River, Mengalong Forest 
Reserve, 5 miles s. w. of Sipitang, 5 ft. alt., 30 
July 1954, G. H. S. Wood & J. Wyatt-Smith 
A4,567 (sing). 
SPECIES FROM CAMBODIA 
SECTION Aero stigma 
Pandanus pendens sp. nov. (sect. Acrostigma) 
Figs. 209, 213^, b 
NOM. VERN.: "rhom cheak.” 
DIAGNOSIS HOLOTYPI: Frutex 7-8 m alta 4 
cm diametro erecta multi-ramosa, cortice brun- 
neo et cum spinis 7-10 mm longis anguste 
conicis, radicibus fulturosis 20-40 cm longis 1 
cm diametro spinosis paucis, foliis 85-92 cm 
longis 2.8-3 cm latis supra obscure viridibus 
infra pallidis et glaucis ligulatis coriaceis in 
sectione mediali cum 28 nervis secundariis in 
quoque dimidio gradatim ex basi in apice subu- 
lato diminuentibus eo 10 cm ex apice 3 mm 
lato, basi amplexicauli et inermi sed ex 4.5-5 
cm marginibus cum dentibus 1.5-2 mm longis 
3-8 mm separatis subulato-serratis, midnervo 
infra ad mediam inermi, in sectione mediali 
marginibus cum serris 0.7-1 mm longis 4-6 
mm separatis adpressis, proxima apicem mar- 
ginibus et midnervo infra cum serrulis 0.2-0. 3 
mm longis 2-4 mm separatis, pedunculo 20-35 
cm longo 10-12 mm diametro obtuse trigono 
folioso-bracteato sed bracteis deciduis, syncarpio 
solitario terminali pendenti 7-10 cm longo 6-8 
cm diametro latiter ellipsoideo, drupis numero- 
sissimis 27-32 mm longis 2.5-3 mm latis 1.5-2 
mm crassis sublinearibus 4-6-angulosis parte l A 
supera libera, pileo laevi anguste ovoideo-conico 
sed aliquis obliquis et proxime inclinatis de- 
ciduis, stylo 2.5-3 mm longo subulato proxime 
arcuato cartilagineo, stigmate 2-2.5 mm longo 
distali sublineari brunneo exlaevi, endocarpio 
in parte 2 A infera cartilagineo brunneo lateribus 
0.1 mm crassis, semine 6-7 mm longo ellipsoi- 
