Page 33 6: Revision of Pandanus, 17. Borneo, Cambodia, Vietnam— St. John 
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deo, mesocarpio apicali viridi mucilaginoso sed 
in sicco cavernoso et 12 mm longo, mesocarpio 
basali in lateribus fibroso sed intra carnoso. 
DIAGNOSIS OF HOLOTYPE: Shrub, 7-8 m tall, 
4 cm in diameter, erect, freely branched above; 
bark brown, with sharp, narrow, conic spines 
7- 10 mm long; prop roots few, 20-40 cm long, 
1 cm in diameter, spiny; leaves 85-92 cm long, 
2.8-3 cm wide, dark green above, pale and 
glaucous below, ligulate, coriaceous, at midsec- 
tion with 28 parallel secondary veins in each 
half, gradually tapering from the base to the 
subulate apex, this 10 cm down 3 mm wide, 
the very base cuneate widened, amplexicaul, 
unarmed, but beginning 4.5-5 cm up the mar- 
gins with teeth 1.5-2 mm long, 3-8 mm apart, 
subulate-serrate; midrib unarmed as far as the 
middle; at midsection the margins with ap- 
pressed serrae 0.7-1 mm long, 4-6 mm apart; 
near the tip the margins and the midrib below 
with serrations 0.2-0. 3 mm long, 2-4 mm apart; 
peduncles 20-35 cm long, 10-12 mm in di- 
ameter, obtusely trigonous, leafy bracted but the 
bracts deciduous before the fruit matures; syn- 
carp single, terminal, pendent, said to become 
”20 cm. long and yellow,” but those seen nearly 
mature (with well-formed seeds) 7-10 cm long, 
6-8 cm in diameter, broad ellipsoid; drupes very 
numerous, 27-32 mm long, 2.5-3 mm wide, 
1.5-2 mm thick, almost linear, 4-6-angled, 
upper Va free; pileus smooth, narrowly ovoid- 
conic, but somewhat oblique and inclined proxi- 
mally, deciduous; style 2.5-3 mm long, subulate, 
proximally arcuate, cartilaginous; stigma distal, 
2-2.5 mm long, almost linear, brown, rough; 
endocarp in lower 36, cartilaginous, brown, the 
walls 0.1 mm thick; seed 6-7 mm long, ellip- 
soid; apical mesocarp continuous, green, muci- 
laginous but on drying shrinking to form a 
cavity 12 mm long; basal mesocarp fibrous along 
periphery, fleshy within. 
HOLOTYPUS: Cambodia, 1 km. s. of Ph. 
Khdat, Vi km. inland, 16 km. w. of Kampot, in 
narrow, wooded stream gulch some 10 m. deep, 
15 m. alt., May 19, I960, Harold St. John 
26,344 ( bish ) . 
DISCUSSION: P. pendens is a member of the 
section Aero stigma, as is its closest relative, P. 
pectinatus Martelli of Sarawak, a species with 
the drupes 27 mm long, 3-4 mm wide, linear 
oblong; endocarp in lower l A, enclosing a conic 
cavity half as long and just above the seed 
cavity. P. pendens St. John of Cambodia has the 
drupes 27-32 mm long, 2.5-3 mm wide, cune- 
ate; endocarp in lower 36, not enclosing a 
superior cavity. 
Local inhabitants reported that the leaves are 
used for plaiting mats, and that after drying 
they are boiled to make a medicine for fevers of 
children. 
The new epithet is the Latin participle pen- 
dens, hanging, given in reference to the posture 
of the fruit. 
section Rykia 
Pandanus cupribasalis sp. nov. (sect. Rykia ) 
Figs. 210, 213 c,d 
NOM. VERN.: ”rhom cheak.” 
DIAGNOSIS HOLOTYPI: Frutex 2-4 m alta 
3-5 cm diametro cum ramis erectis pluribus 
plerumque simplicibus, cortice brunneo laevi 
vel cum radicillis spinosis paucis, radicibus ful- 
turosis 30-60 cm longis 1 cm diametro sub- 
laevibus sed cum radicillis conicis paucis re- 
motis, foliis multis 1.48 m longis 3.8-4 cm 
latis supra obscure viridibus infra viridibus late 
sulcatis et subplicatis ligulatis sed in 20 cm 
ultimis in apice trigono subulato abrupte di- 
minuentibus eo in puncto 10 cm ex apice 1.6 
mm lato, in sectione mediali cum 31-34 nervis 
secundariis parallelis in quoque latere et ad 
apicem infra cum nervis tertiis transversis evi- 
dentis et reticulis anguste oblongis formanti- 
bus, basi amplexicauli inermi intense cupracea 
et ex 15-25 cm marginibus cum spinis 4-5 mm 
longis 12-18 mm separatis crassiter subulatis 
arcuatis adscendentibus pallidis excepta apicibus 
Fig. 207. Pandanus trigonus St. John, from holotype. a, Syncarp, X 1; h, drupe, lateral view, XI ; G drupe, 
longitudinal median section, XI \ d, drupe, lateral view, X 4; e, drupe, longitudinal median section, X 4; 
/, drupe, apical view, X 4; g, drupe, apical view, X 10; h, leaf base, lower side, X 1 \ i, leaf middle, lower side, 
X 1; j, secondary and tertiary veins, lower side, at middle, X 4; k, leaf apex, lower side, X L 
