Page 338: Revision of Pandanus, 17. Borneo, Cambodia, Vietnam — St. John 
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brunneis, midnervo infra ex 25 cm cum aculeis 
5-7 mm longis 18-22 mm separatis crassiter 
subulatis reflexis, in sectione mediali marginibus 
cum aculeis 1.8-2 mm longis 10-25 mm sepa- 
ratis subulatis adpressis adscendentibus, mid- 
nervo infra subinermi, proxima apicem margini- 
bus et midnervo infra cum serrulis 0. 3-0.6 mm 
longis 2-6 mm separatis, pedunculo 50 cm 
longo in apice 13 mm diametro infra graciliori 
folioso-braceato cernuo terminali, syncarpio 
unico 16.5 cm longo 8 cm diametro oblongo- 
ellipsoideo 3-laterato cum circa 540 drupis et 
superficie spinoso ex stylis salientibus, drupis 
25-30 mm longis 7-10 mm latis 5-9 mm crassis 
cuneato-oblanceoloideis viridibus sed apparente 
lutescentibus laevibus 6-lateratis lateribus planis 
parte Va supera libera, pileo pyramidali-semi- 
orbiculari, stylo 3-4 mm longo valido osseoso 
crasso lucido corniformi brunneo in 40° adscen- 
denti latiter bifido ( rare trifido, et eis apicalibus 
simplicibus ) , stigmate 3 mm longo plerumque 
latiter ovato et in apice bifido brunneo papil- 
lose, endocarpio minime submediali aurantiaco- 
brunneo lateribus 2 mm crassis, semine 8-9 mm 
longo 3 mm diametro anguste ellipsoideo; rneso- 
carpio apicali cavernoso et cum fibris longitudi- 
nalibus paucis et membranis medullosis, meso- 
carpio basali fibroso et carnoso. 
DESCRIPTION OF ALL SPECIMENS EXAMINED: 
Shrub 2-4 m tall, 3-5 cm in diameter, with 
several erect, mostly simple stems; bark brown, 
smooth, or with a few latent rootlets; prop roots 
few, 30-60 cm long, 1 cm in diameter, smooth, 
except for a few remote conic rootlets; leaves 
in terminal plumes, numerous, 1.48-3.6 m long, 
3.8-4 cm wide, above dark green, below green, 
with a broad central groove and gently down 
curved outer quarters, ligulate, then about 20 
cm from the tip abruptly narrowed into a subu- 
late trigonous tip which 10 cm down is 1.6 mm 
wide, at midsection with 3 1-34 parallel second- 
ary veins in each half, towards the apex the 
lower side with evident tertiary cross veins 
making a reticulum of narrow oblong meshes, 
the amplexicaul, hidden base unarmed, bright 
copper-orange, beginning at 15-25 cm up the 
margins with spines 4-5 mm long, 12-18 mm 
apart, heavy subulate, arcuate, ascending, pale, 
but the tip brownish; the midrib below, begin- 
ning 15-25 cm from the base with stout subulate 
spines 5-7 mm long, 18-22 mm apart, reflexed; 
at midsection the margins with prickles 1.8-2 
mm long, 10-25 mm apart, subulate, appressed 
ascending, the nearby midrib below almost 
unarmed; near the apex the margins and midrib 
below with serrulations 0. 3-0.6 mm long, 2-6 
mm apart; peduncle 50 cm long, at apex 13 mm 
in diameter, tapering and more slender down- 
wards, leafy bracted, nodding, terminal; syncarp 
single, 16.5 cm long, 8 cm in diameter, oblong- 
ellipsoid, distinctly 3 -sided, with about 540 
drupes, the surface thorny from the projecting 
styles; drupes 25-30 mm long, 7-10 mm wide, 
5-9 mm thick, cuneate-oblanceoloid, still green, 
apparently becoming yellowish, the surface 
smooth, 6-sided, the sides plane, upper Va free; 
pileus pyramidal-semiorbicular; stylar beak 3-4 
mm long, stout, bony, shining, brownish, as- 
cending at about 40°, almost all widely bifid, 
but rarely one is trifid, and the apical ones 1- 
dentate; stigma 3 mm long, mostly broad ovate 
and bifid at apex, brown, papillose; endocarp 
slightly submedian, orange-brown, the lateral 
walls 2 mm thick; seed cavity 8-9 mm long, 3 
mm in diameter, narrowly ellipsoid, traversed 
by longitudinal fibers; apical mesocarp cavern- 
ous, with a few longitudinal fibers and medullary 
membranes; basal mesocarp fibrous and fleshy. 
HOLOTYPUS: Cambodia, Chaine de l’Elephant, 
on the mountain grade 12 km. below Le Bokor, 
dense moist fern forest, with Ficus , Homalan- 
thus, and Hicriopteris, 1,000 m. alt., May 20, 
I960, H. St. John 26,346 (bish). 
specimens examined: Cambodia, Bokor, 
common in ever green forest, 900 m. alt., 21 
Feb. I960, Tem Smitinand 6,334 (bkf). 
DISCUSSION: P. cupribasalis is a member of 
the section Rykia, as is its closest relative, P. 
furcatellus Martelli, of Indochina, a species with 
Fig. 208. Pandanus tectorius Soland. ex Warb. var. horneensis Martelli, from Wood & Wyatt-Smith A4,567. 
a, Phalange, lateral view, X 1; ^ phalange, longitudinal median section, X C o, phalange, apical view, X 1; 
d, carpel apex and stigma, oblique view, X 4; e, leaf base, lower side, X 1; f, leaf middle, lower side, X 1; 
g, leaf apex, lower side, X 1- 
