Page 340: Revision of Pandanus, 17 . Borneo, Cambodia, Vietnam — St. John 
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the syncarp covered with bracts, solitary, erect; 
styles ovate, acute; endocarp 1.5 cm long, the 
lateral walls 0.8 mm thick; seed cavity truncate 
at apex, 6 mm in diameter; leaves 8 cm wide, 
near the base the margins with spines 6 mm 
long, 14-23 mm apart, stout arcuate subulate, 
the tips brownish, on the midrib below the 
spines 7-8 mm long, 22-40 mm apart, similar 
but reflexed and the tips slender. P. cupribasalis 
has the syncarp exposed, solitary, nodding; styles 
lanceolate, acute; endocarp 2 cm long, the lateral 
walls 2 mm thick; seed cavity ellipsoid, 4 mm 
in diameter; leaves 3.8-4 cm wide, near the base 
the margins with spines 4-5 mm long, 12-18 
mm apart, those of the nearby midrib below 5-7 
mm long, gradually narrowed to a long subulate 
apex. 
The new epithet is formed from the Latin 
cuprum, copper; basalis, of the base, and is 
given in allusion to the strikingly copper-colored 
leaf bases. 
SPECIES FROM VIETNAM 
SECTION Rykia 
Pandanus depressus sp. nov. (sect. Rykia) 
Figs. 211, 21 Aa 
DIAGNOSIS HOLOTYPI: Caulibus foliisque non 
conservatis (collector dicet ad P. horizontalis 
simulantibus, id est caule 1.5 m alto 4-5 cm 
diametro cum radicillis adpresse adscendentibus 
paucis, foliis 2.72 m longis 3.8 cm latis olivaceis 
2-plicatis ligulatis abrupte in apice subulato 
caudato contractis, marginibus et midnervo infra 
cum aculeis validis), inflorescentia foeminea 
terminali cum syncarpio solitario, pedunculo 17 
cm vel ultra longo 12 mm diametro 3-laterato 
folioso-bracteato, syncarpio 15 cm longo 8 cm 
diametro cylindrico cum apicibus rotundatis et 
cum circa 816 drupis submaturis viridibus, 
drupis in lineis subverticalibus eis basalibus et 
apicalibus minime latioribus, drupis 30-32 mm 
longis 8-12 mm latis 6-10 mm crassis anguste 
cuneatis 6-angulosis parte Vs supera libera cor- 
pore 26-28 mm longo, pileo cum basi 5-6 mm 
alto valde depresse rotundato-pyramidali laevi 
glauco, stylo terminali validi corniformi sube- 
recto vel minime proxime curvato eis apicalibus 
valde curvato 1.5-2 mm alto furcis 2-3 mm lon- 
gis horizontali-divergentibus osseosis acutis pau- 
cis cum lobis lateralibus binis proximalibus, stig- 
matibus 1.5-2. 5 mm longis lanceolatis brunneis 
papillosis non ad apices extends semiproximali- 
bus et terminalibus, endocarpio submediali os- 
seoso brunneo et cum apice mediali acuminate 
lateribus 1 mm crassis, semine 8-9 mm longo 4 
mm diametro obovoideo, mesocarpio apicali 
medulloso molli albo exfibroso, mesocarpio ba- 
sali in lateribus fibroso in centro carnoso. 
DIAGNOSIS OF HOLOTYPE: Stem and leaves 
not collected (but said by the collector to be 
similar to his no. 2,519, P. horizontalis; that is, 
stem 1.5 m tall, 4-5 cm in diameter, with a few 
appressed ascending rootlets; leaves 2.72 m long, 
3.8 cm wide, olive green, 2 -pleated, ligulate, 
abruptly contracted to a caudate, subulate apex, 
the margins and midrib below with stout 
prickles); pistillate inflorescence terminal, bear- 
ing one syncarp; peduncle more than 17 cm long, 
12 mm in diameter, 3-sided, leafy bracted; syn- 
carp 15 cm long, 8 cm in diameter, cylindric 
with rounded ends, bearing about 816 drupes, 
somewhat immature, green, but with hard en- 
docarp and well-formed seeds, probably nearly 
full size; drupes arranged in nearly vertical rows, 
the basal and apical drupes slightly broader, but 
not significantly so; drupes 30-32 mm long, 8- 
12 mm wide, 6-10 mm thick, narrowly wedge- 
shaped, 6-angled, upper Vs free, the body 26-28 
mm long; pileus with its base 5-6 mm high, 
much depressed, rounded pyramidal, smooth, 
glaucous; style terminal, stout, hornlike, suberect 
or slightly curved proximally, the apical ones 
definitely so, 1.5-2 mm high, the two forks 
lanceolate, 2-3 mm long, spreading horizontally, 
hard, sharp, mucronate and unpleasant to handle, 
sometimes with 2 small proximal lateral lobes; 
stigmas 1.5-2. 5 mm long, lanceolate, brown, 
papillose, not reaching to the tip, semiproximal 
Fig. 209. Pandanus pendens St. John, from holotype. a, Drupe, lateral view, X 1; ^ drupe, longitudinal 
median section, X 1; c, drupe, lateral view, X 4; d, e, drupe, longitudinal median section, X 4; f, carpel apex 
and stigma, distal view, X 15; g, leaf base, lower side, X 1 ^ leaf middle, lower side, X 1; 7 leaf apex, 
lower side, X 1. 
