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Page 342: Revision of Pandanus, 17. Borneo, Cambodia, Vietnam — S t. John 
and terminal; endocarp submedian, bony, brown, 
with an apical subulate prolongation, the walls 
1 mm thick; seed 8-9 mm long, 4 mm in diame- 
ter, obovoid; apical mesocarp soft white, pithy, 
without fibers; basal mesocarp fibrous up the 
periphery, fleshy within. 
holotypus: Vietnam, Hon Thom, lie S. de 
Phu quoc, bord de ruisseau, 30-50 m. de rivage, 
a 1’ombre des arbres, 7 janvier 1961, Pham 
Hoang Ho 2,530 (bish). 
DISCUSSION: P. depressus is a member of the 
section Rykia, as is its closest relative, P. hori- 
zontalis St. John, a species from the same island, 
but with syncarp broadly ellipsoid, 10.5 cm long; 
terminal and subterminal drupes the larger; 
pileus base conic; styles often with 2-4 second- 
ary lobes; stigmas terminal; endocarp in lower 
14, the walls 0.1 mm thick; seed 2.5 mm in 
diameter. P. depressus has the syncarp cylindric, 
15 cm long; drupes all about the same size; 
pileus base much depressed rounded pyramidal; 
styles sometimes with 2 secondary lobes; stigmas 
semiproximal and terminal; endocarp subme- 
dian, the walls 1 mm thick; and the seed 4 mm 
in diameter. 
The new epithet is the Latin adjective de- 
pressus, depressed, and is given in reference to 
the depressed rounded pyramidal pileus base. 
Pandanus horizontalis sp. nov. (sect. Rykia ) 
Figs. 212, 214b 
diagnosis HOLOTYPI: Caulibus 1.5 m altis 
4-5 cm diametro lucidis cum radicillis adpresse 
adscendentibus paucis, foliis 2.72 m longis 3.8 
cm latis infra olivaceis supra obscure ita et pro- 
funde sulcatis 2-plicatis, in sectione mediali cum 
33 nervis secundariis parallelis in quoque latere, 
proxima apicem nervis tertiis transversis evi- 
dentis et reticulis oblongis formantibus, lamina 
ligulata in apice 20 cm longo trigono subulato 
caudato abrupte contracta eo 10 cm ex apice 1 
mm lato, basi amplexicauli inermi exlucida cu- 
pracea et ex 10-14 cm marginibus cum aculeis 
2-4 mm longis 9-18 mm separatis crassiter 
subulatis rectis adscendentibus apicibus brunneis, 
midnervo infra ex 8 cm cum spinis 3-4 mm 
longis 17-35 mm separatis conicis reflexis brun- 
neis, in sectione mediali marginibus cum aculeis 
2- 3 mm longis 15-25 mm separatis crassiter su- 
bulatis adpresse adscendentibus albis sed apicibus 
brunneis, midnervo infra cum spinis reflexis eis 
basalibus simulantibus sed 6-8 cm separatis, in 
sectione apicali diminuenti marginibus et mid- 
nervo infra cum serris 0.5-1 mm longis 1.5-5 
mm separatis, in apice subulato marginibus cum 
serrulis 0.3-0. 5 mm longis 1.5-3 mm separatis 
translucentis sed apicibus brunneis, inflorescen- 
tia foeminea terminali cernua cum syncarpio 
unico, pedunculo 45 cm longo 1.8 cm diametro 
3- laterato folioso-bracteato, bracteis interioribus 
15 cm longis 4.4 cm latis lanceolatis marginibus 
et midnervo infra serratis, syncarpio 10.5 cm 
longo, 7.8 cm diametro latiter ellipsoideo cum 
circa 720 drupis, eis submaturis viridibus lute- 
scentibus eis lateralibus 26-27 mm longis 6-9 
mm latis 5-7 mm crassis anguste oblanceoloideis 
parte Vs supera libera illis terminalibus et sub- 
terminalibus 30-31 mm longis 9-11 mm latis 
7-10 mm crassis anguste oblanceoloideis 6- 
angulosis corpore 25-28 mm longo, pileo cum 
basi 5-6 mm alta rotundato-conica 6-angulosa 
laevi viridi subglauca, stylo druparum lateralium 
erecto solido bifurcato furcis lanceolatis acerosis 
in angulis rectis divergentibus, stigmatibus lan- 
ceolatis brunneis papillosis terminalibus sed 
lobatis in stylis multis cum lobulis secundariis, 
basi 1-2 mm longa lobis principalibus 2. 5-3. 5 
mm longis, endocarpio in parte Vs infera brun- 
neo ellipsoideo acuminato lateribus 0.1 mm 
crassis cartilagineis, semine 12 mm longo 2.5 
mm diametro ellipsoideo, mesocarpio apicali 
medulloso humido sine fibris, mesocarpio basali 
3 mm longo in lateribus fibrosis in centro 
carnoso. 
DIAGNOSIS OF HOLOTYPE: Stem 1.5 m tall, 
4- 5 cm in diameter, shining, with a few ap- 
pressed ascending rootlets; leaves 2.72 m long, 
FIG. 210. Pandanus cupribasalis St. John, from holotype. a, Drupe, lateral view, X L b, drupe, longitu- 
dinal median section, X 1 \ c, d, f, h, i, typical lateral drupes, apical view, X 1; ^ apex and stigma of apical 
drupe, proximal view, X 4; j, apex and stigma of typical lateral drupe, proximal view, X 4; k, apex and 
stigma of rare 3-fid lateral drupe, apical view, X 4; /, leaf base, lower side, X 1 leaf middle, lower side, 
X 1; #, leaf apex, lower side, X 1; o, veins, lower side, towards apex, X 4. 
