Page 344: Revision of Pandanus, 17. Borneo, Cambodia, Vietnam — St. John 
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3.8 cm wide nearly throughout, olive green be- 
low, darkly so above with a deep furrow over 
the midrib, 2 -pleated, at midsection with 33 
secondary parallel veins in each side, and near 
the tip tertiary cross veins visible, forming ob- 
long meshes; blade ligulate, abruptly contracted 
to a 20 cm subulate, trigonous, caudate apex, 
this 10 cm down 1 mm wide, the base amplexi- 
caul, unarmed, dull coppery, beginning at 10- 
14 cm the margins with prickles 2-4 mm long, 
9-18 mm apart, heavy subulate, straight, ascend- 
ing, brown-tipped; the midrib below beginning 
at 8 cm with thorns 3-4 mm long, 17-35 mm 
apart, conical, reflexed, brown; at midsection the 
margins with prickles 2-3 mm long, 15-25 mm 
apart, thick subulate, appressed ascending, white 
below, with a brown tip; the midrib below with 
reflexed spines like the basal ones but 6-8 cm 
apart; where the blade narrows the margins and 
midrib below with serrations 0.5-1 mm long, 
1.5-5 mm apart; on the subulate apex the 
margins with serrulations 0.3-0. 5 mm long, 1.5- 
3 mm apart, brown-tipped, but the rest clear 
and translucent; pistillate inflorescence terminal, 
cernuous, with a single syncarp; peduncle 45 
cm long, 1.8 cm in diameter, 3 -sided, leafy 
bracted; the inner bracts 15 cm long, 4.4 cm 
wide, lanceolate, serrate on margins and midrib 
below; syncarp 10.5 cm long, 7.8 cm in diameter, 
broadly ellipsoid, bearing about 720 drupes, 
these immature, green, but some turning yellow- 
ish, the lateral ones 26-27 mm long, 6-9 mm 
wide, 5-7 mm thick, narrowly oblanceoloid, 
upper Vs free, the terminal and subterminal 
ones 30-31 mm long, 9-11 mm wide, 7-10 mm 
thick, narrowly oblanceoloid, 6-angled, the body 
25-28 mm long; pileus with its base 5-6 mm 
high, rounded conic, 6-angled, smooth, green, 
slightly glaucous; style of lateral drupes erect, 
stout, bifurcate, the forks lanceolate, spreading 
horizontally, acerose tipped; stigmas lanceolate, 
brown, papillose, terminal, but on the many 
styles with 2-4 short proximal secondary lobes, 
running over them also, style base 1-2 mm 
long, main lobes 2. 5-3. 5 long; endocarp in lower 
Vi, brown, ellipsoid, with an apical subulate 
extension to the style, the walls cartilaginous, 
0.1 mm thick; seed 12 mm long, 2.5 mm in 
diameter, ellipsoid; apical mesocarp a moist pith, 
without fibers; basal mesocarp 3 mm long, 
fibrous up the periphery, fleshy within. 
HOLOTYPUS: Vietnam, lie Hon Tho’m, s. of 
lie Phu quoc, 7 avril 1961, Pham Hoang Ho 
2,519 (bish). 
DISCUSSION: P. horizontalis is a member of 
the section Rykia. It has the hornlike, bifurcate 
style of that section and on the terminal and 
subterminal drupes the style is bent proximally 
and the stigmas are proximal as is normal. On 
all the lateral drupes, which are very numerous, 
the style is erect, with its forks extending later- 
ally at right angles, and the stigmas are terminal. 
This is exceptional, but from the fact that the 
style is hornlike, and bifurcate, and that some 
of them incline in the proximal direction, it is 
concluded that this new species really belongs 
in the section Rykia. 
No close relative of this new species has been 
described. The closest is an undescribed species 
from Thailand, Chanburi, Tern Smitinand 4,054, 
but this has smaller, broader drupes, etc. 
The new epithet is the Latin adjective hori- 
zontalis, horizontal, and is given in reference to 
the horizontal forks of the style and the stigmas. 
LOCALITY RECORDS OF PANDANUS IN VIETNAM 
Pandanus ceratostigma Martelli 
Region de Long cau, Col de Tuy lu’at, at km. 
1,267, male, 6 fevrier 1961, P. H. Ho 2,605 
(BISH, univ. SAIGON). Also his no. 2,610. 
Col de Dio Ca( Varella) , pres du poste mili- 
taire, km. 1,364, 6 fevrier 1961, Pham Hoang 
Ho 2,604 (BISH, UNIV. SAIGON). 
Pandanus integriapicis St. John 
Prov. de Phu yen, Anthanh (Tuyan), km. 
FIG. 211. Pandanus depressus St. John, from holotype. a, Drupe, lateral view, X 1; ^ drupe, longitudinal 
median section, X 1; c, d, e, f, g, h, i, drupe, apical view, X 1 \ j, drupe, lateral view, X 4; k, drupe, longi- 
tudinal median section, X 4; l, drupe apex and stigma, apical view, X 4; m, drupe apex and stigma, oblique 
distal view, X 4. 
