Revision of the Genus Pandanus Stickman, Part 19 
Additional Malayan Species of Pandanus 
Harold St. John 
Most of the Malayan species of Pandanus 
have already been treated by the writer in parts 
11, 14, and 15 of this revision. Here are pre- 
sented descriptions of five of the remaining 
species. 
Pandanus elostigma Martelli, Soc Bot. I tal., Bui. 
n. s. 11 (2): 302-303, 1904; Webbia 4(1): 
13, 1913; 4(2) :t. 30, fig. 9, 1914; Ridley, 
FI. Malay Benin. 5:79, 1925. (sect. Acro- 
stigma ) . 
Fig. 219 
NOM. VERN. : "shidayen masing.” 
DIAGNOSIS OF LECTOTYPE: Apparently a low 
shrub, growing in clusters of as many as 20; 
leaves 1-2.24 m long, 18.5-22 mm wide, thick 
firm chartaceous, above dark bluish green, be- 
low bluish green, 1 -ribbed, 2 -pleated, the central 
furrow narrow, in section M-shaped, at midsec- 
tion with 19-22 parallel secondary veins in each 
half, these prominent throughout, tertiary cross 
veins visible l A way from the base and in outer 
half conspicuous, more or less transverse and 
forming short oblong meshes, the blade ligulate 
but in upper 1/6 gradually narrowing to a 
trigonous subulate apex about 5 cm long, 1.5-2 
mm wide, the base unarmed, but beginning 13- 
lb cm up the margins with prickles 0,6-0. 9 
mm long, 3-5 mm apart, heavy subulate, as- 
cending, pale; the midrib below with a few 
prickles beginning at 10.5 cm up and 1-1.7 mm 
long, 4-11 mm apart, conic subulate, reflexed; 
at midsection the margins with delicate prickles 
0.4-0. 5 mm long, 3-5 mm apart, subulate, 
closely appressed; the nearby midrib unarmed; 
on the subulate apex the margins with subulate 
tipped serrulations 0. 3-0.4 mm long, 1-2 mm 
apart; the midrib below with similar serrulations 
1.5-7 mm apart; syncarp 8-12.5 cm long, 5-7.5 
1 B. P. Bishop Museum, Honolulu, Hawaii 96819, 
U.S.A. Manuscript received August 13, 1962. 
subglobose; drupes 23-26 mm long, 5-6 mm 
wide and thick, fusiform, the body 11-13 mm 
long, oblanceoloid, fleshy but when dry the apex 
shrunken and much smaller than the flaring base 
of the pileus; pileus 1.5-2 mm high, smooth, 
the base discoid, firm, but when dried flaring 
and undulate; style 9-10 mm long, thick 
subulate, mostly proximally curved, the lower 
half with 4-6 sharp angles but the inter- 
vals flat or furrowed, the upper half terete; 
stigma 6-7 mm long, linear, distal, brown, papil- 
lose, running to the tip; endocarp centering in 
lower 2/7, cuneate the walls 0.1 mm thick, pale 
brown, except near the apex, formed principally 
of the heavy, longitudinal, marginal fibers, these 
connected by a thin tissue; apical mesocarp 5 
mm long, ellipsoid from a truncate base, con- 
taining only the white membranes of an aer- 
enchyma; basal mesocarp fleshy and fibrous. 
LECTOTYPE: Malay Peninsula, Perak, Larut, 
in large clusters on rocky soil, 300 to 800 ft alt, 
Sept. 1884, Dr. King’s Collector 6,559 (cal)! 
Isotype (SING)! 
SPECIMENS EXAMINED: Malaya, Perak, Larut, j; 
open ground, 300 ft alt 1883, Dr. King’s Col- 
lector 3,758 (cal, sing) This is also in (fi) 
but by error as 3,755 . Malacca, Gaong dalam 
Ayer Panas, Oct. 1893, J. S. Goodenough 1,535 
(CAL). 
DISCUSSION: As Martelli originally described 
this species from two collections, one is here 
chosen as lectotype. 
The illustrations of two drupes published by 
Martelli in 1914 show well the remarkable flar- 
ing base of the pileus. Our illustration, Figure 
219 b, e, shows a dried drupe similar to his, but 
our Figure 219 a, d, shows one after boiling. In 
this one the lower pileus flange has rounded out 
224 
