229 
Page 359: Revision of Pandanus, 19- Malaya — S t. John 
margins with prickles 0.5-1 mm long, 4-11 
mm apart, subulate, flat appressed ascending; 
along the outer third on the upper surface the 
two lateral pleats with subulate tipped serra- 
tions 0.3— 0.5 mm long, 3-10 mm apart; on the 
caudate apex the margins and midrib below 
with serrations 0.3-0. 5 mm long, those on the 
margins 1-2.5 mm apart, those on the midrib 
below 2-4 mm apart; pistillate inflorescence 
terminal, the tip pendent, bearing 1 syncarp 
(or rarely with 2-5 lateral, smaller, secondary 
ones); peduncle 20-52 cm long, 4-9 mm in 
diameter, 3 -sided, bracteate; syncarp 12-23 cm 
long, 2. 5-5. 5 cm in diameter, cylindric, with 
obtuse ends, bearing 400-1,728 drupes, these 
11-15 mm long, or following the curve of the 
style 13-18 mm long, 3-4 mm wide and thick, 
oblong-oblanceolate, with a subulate apex, 
4-6-angled, the body 9-14 mm long; pileus 
2.5-4 mm high, obliquely conic, smooth, usu- 
ally obscurely angled; style 3. 5-5. 5 mm long, 
subulate, sharply arcuate proximally, rigid; 
stigma 2-4 mm long, linear, distal, brown, 
papillose; endocarp centering in lower l A, the 
walls 0.1 mm thick, stramineous; apical meso- 
carp semiorbicular, filled with parenchyma; 
basal mesocarp sparse, fibrous and fleshy. 
DESCRIPTION OF STAMINATE PLANT ( Ridley 
12,143 ): Plant with similar leaves; inflores- 
cence 25 cm long; the bracts colored (white ?), 
subherbaceous; lowest floral bract with a col- 
ored base 6 cm long and a foliaceous prolonga- 
tion 15 cm long, 18 mm wide, ligulate, taper- 
ing gradually into a subulate apex; median 
floral bract 9 cm long, 2 cm wide, lanceolate, 
unarmed elsewhere, but the tip with margins 
and midrib below with yellow serrulations 0.2 
mm long, crowded; the 7-8 spikes 3-6 cm 
long, 10-12 mm in diameter, dense, having the 
odor of crushed leaves of P. amaryllifolius; 
; stamens separate, attached directly to the rha- 
chis, filaments 0.2— 0.3 mm long; anther 3.5- 
4 mm long, linear, bearing a subulate pro- 
longation of the connective 0. 5-0.6 mm long. 
HOLOTYPE: Malaya, Perak, Taiping Hill, 
1,000 ft elev, 17 Feb. 1907, Md. Haniff & Md. 
Nur 2,393 (sing)! Isotype (k) ! 
SPECIMENS EXAMINED: Malaya, Malacca, 
j Gaudichaud (p), holotype of Fisquetia ornata 
Gaud; also in (g), isotype; and (FI), clasto- 
type; Malacca, without . collector (Fl); Perak, 
Larut, Gunong Boobo, 2,500-3,000 ft alt, 
March 1885, H. Kunstler 7,374 (fi); Sungei 
Penang Rd., 1,000 ft alt, Sept. 1887, C. Curtis 
1,220 (K); Gunong Bintang, Kedah-Perak 
Boundary, June 1917, from Fed. Malay States 
Mus. (k); Tanjong Malim, 3 July 1924, 7. H. 
Bur kill & Md. Haniff 1 4,000 ( K ) ; Kedah, Koh 
Moi Forest Res., 5 April 1938, Kiah 33,190 
(k); Penang, Government Hill, Ridley 2,272, 
type of P. ornatus f . spicatus Martelli (FI, K ) ; 
Singapore, Bukit Timah, 1894, H. N. Ridley 
6,287 (k); Johore, top of St. George, Gunong 
Pulai, Dec. 1905, Ridley 12,143 (k, sing), 
staminate; Trengganu, S. Trenganu Ferry, 26 
April 1937, E. f. H. Corner 33,473 (k) ; Negri 
Sembilan, Tampin Hill, June 16, 1917, Ridley 
(k); Singapore, cult, in Bot. Garden, 1902, 
Ridley (Fl) ; Johore, 2 miles N. E. of Kampong 
Sungei Sedili Besar, edge of forest at sea shore, 
2 June 1962, /. Sinclair 10,703 (e); Kedah, 
Ulu Sungei Terenas, 20.3.1938, Sow 46,187 
(kep); Selangor, Ulu Gombak, 17.6.193 1, C. F. 
Symington 24,334 (kep); Kedah, Kedah Peak, 
9.5.1938, Symington 46,937 (kep); ditto, very 
common near top, 3,700 ft alt, 18/5/1957, 
Wyatt-Smith 79,292 (kep). 
Thailand: Bachaw, Pattani, evergreen forest, 
600 m alt, July 14, 1923, A. F. G. Kerr 7,191 
(BK). 
DISTRIBUTION : Throughout Malaya, near 
streams in lowlands and in forests on hills up 
to 3,000 ft alt; also in southern Thailand. 
Discussion: P. recurvatus is a member of the 
section Rykia, as is its closest relative, P. mono- 
theca Martelli, of Malaya, a species with the 
peduncle erect; syncarp 5 cm long, ovate- 
ellipsoid; drupes 10-13 mm long, obovoid or 
narrowly so, the body 7-9 mm long; pileus 
broadly ovoid; seed 4-5 mm long, obconic; 
leaves 0.7-1. 1 m long, at midsection with 18 
parallel secondary veins in each half, beginning 
5 cm up from the base the margins with prickles 
2-2.5 mm long, 2-5 mm apart, subulate sig- 
moid, and the lower ones recurving. P. recurva- 
tus has the peduncle recurved; syncarp 12-23 
cm long, cylindric; drupes 13-18 mm long, 
oblong-oblanceoloid, the body 9—14 mm long; 
