462 
PACIFIC SCIENCE, Vol. XIX, October 1965 
holotype: U. S. Nat. Mus. Helm. Coll., S.Y. 
No. 20. 
DESCRIPTION (based on 13 mature whole 
mounted specimens ) : Body elongate fusiform, 
with blunt extremities, 3.75-11 mm in length 
with maximum width of 0.8-2. 3 mm in ovario- 
testicular region where the lateral margins are 
more or less irregularly crenulated; cuticle thick, 
beset throughout with minute spines. Oral 
sucker ventroterminal, 0.23-0.5 X 0.23-0.5 mm; 
prepharynx 0.13-0.55 mm long; pharynx spher- 
ical, 0.18-0.32 mm in diameter; esophagus 30- 
200 ,u long; ceca simple and wide anteriorly, 
but may be diverticulate on outer side in hind- 
body, opening outside separately, one on each 
side of excretory pore, at posterior extremity. 
Acetabulum 0.28-0.64 mm in diameter, situ- 
ated at or near posterior end of anterior third 
of body. 
Testes deeply Iobed, 0.18-1.3 X 0.25-1.2 
mm, tandem; anterior testis usually 5-lobed, at 
posterior end of middle third of body, separated 
from ovary as well as from posterior testis by 
vitellaria; posterior testis usually 6-lobed, at 
anterior part of posterior third of body. Sem- 
inal vesicle tubular, 50-170 /x wide, closely 
winding posterodorsal to acetabulum in the 
fully gravid type, but extending farther back- 
ward in young individuals, a little to left of 
median line, with its anterior portion sur- 
rounded by prostate cells, whose ducts enter 
the cirrus pouch at its posterior end to dis- 
charge into the pars prostatica enclosed in the 
pouch; the anterior end of the seminal vesicle 
penetrates the posterior end of the cirrus 
pouch, and forms a definite, distally swollen, 
internal seminal vesicle which leads into the 
pars prostatica with a constriction or demarca- 
tion between. Pars prostatica muscular, bulbous, 
with its distal end differentiated into a short 
cirrus. Cirrus pouch saccular, thin-walled, 0.2- 
0.6 X 0.09-0.4 mm, extending obliquely or 
transversely from anterosinistral edge of acetab- 
ulum to genital pore, containing distal end of 
external seminal vesicle, internal seminal vesi- 
cle, very strongly developed pars prostatica, nu- 
merous prostatic ducts coming from prostate 
cells surrounding distal portion of external sem- 
inal vesicle, and a short stout cirrus. Genital 
atrium muscular, lined with thick cuticle, open- 
ing sinistrosubmarginally at pre-acetabular level. 
Ovary subglobular to longitudinally elongated 
oval, 0.06-0.56 X 0.06-0.4 mm, at or behind 
equatorial level, slightly to right of median line. 
Germiduct arising from left side of ovary, 
strongly constricted at its origin, giving off 
Laurer’s canal just before joining seminal re- 
ceptacle; seminal receptacle large, 0.2-0.5 X 
0.15-0.4 mm in fully mature specimens, situ- 
ated to left of ovary, a little in front of anterior 
testis. Laurer’s canal opening outside dorsal or 
immediately sinistral to seminal receptacle. 
Uterus tightly coiled in intercecal field between 
ovary and acetabulum; metraterm running 
alongside male terminalia. Eggs oval, 44-54 X 
27-37 n in life. Vitelline follicles comparatively 
small, circumcecal, commencing at level of 
posterior end of acetabulum on the right, but 
a little more posteriorly on the left, intruding 
into space between ovary and anterior testis as 
well as into intertesticular space without meet- 
ing in median line, but almost confluent in 
posttesticular area; vitelline reservoir oval, 85 ft 
wide in the type, up to 0.28 X 0.18 mm, situ- 
ated ventrally between ovary and seminal re- 
ceptacle. Excretory vesicle Y-shaped; its stem 
with numerous lateral diverticles reaching to 
intestinal limbs or overlapping them; in post- 
testicular area these lateral diverticles are sim- 
ple, but the remaining are irregularly subdi- 
vided dendritically. Between the acetabulum and 
the ovary the diverticles anastomose with one 
another in median line in mature specimens, so 
that the primary median stem recognized in 
immature specimens is now replaced by anas- 
tomosing diverticles. Immediately behind the 
intestinal bifurcation the median stem divides 
into two lateral arms, each of which gives off 
an inner secondary branch communicating an- 
teriorly with the main lateral branch of its own 
side; excretory pore terminal, not forming 
cloaca. 
DISCUSSION: This new genus closely resem- 
bles Allolepidapedon Yamaguti, 1940, but the 
general body shape, the peculiar structure of 
the excretory vesicle, and the presence of two 
separate anal openings instead of the cloaca 
prevent its being identified with the latter 
genus. It is defined as follows: 
