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PACIFIC SCIENCE, Vol. XIX, October 1965 
thickening upwards and near the apex 4 mm in 
diameter; syncarp 3.5 cm long, 2.5 cm in diam- 
eter, oblong-ellipsoid, bearing about 720 drupes, 
these 11—12 mm long, 2 mm wide and thick, 
5-6-angled, the body 7-8 mm long, oblong- 
oblanceoloid; pileus with its base 1.5-2 mm 
high, semiellipsoid, the surface smooth or 
slightly papillose; style 5-6 mm long, slender 
subulate, bent at base and sharply reflexed, the 
upper drupes with nearly terminal styles, the 
lower ones with styles lateral in attachment; 
stigma 4-5 mm long, linear, proximal, brown, 
papillose, running almost to the stigma tip; en- 
docarp in lower V5, the walls 0.07 mm thick, 
cartilaginous, stramineous; seed 2.5 mm long, 
ellipsoid; apical mesocarp sparse, pithy; basal 
mesocarp fibrous up the margins, fleshy within. 
HOLOTYPUS: India, Assam, Kambang, plains, 
May 1888, L. Wray, Jr. 1,939 (sing). Isotype 
(CAL). 
DISCUSSION: P. assamensis is a member of 
section Rykia, as is the most closely related spe- 
cies, P. Scortechinii Martelli, of Perak, Malaya. 
The new holotype was included among the spec- 
imens cited by Martelli when he published the 
latter species. Later, he typified his species by a 
collection from Perak, Malaya, and omitted the 
collection from Assam. One of these is made the 
basis of the species here described as new. P. 
Scortechinii has the style 4—5 mm long, subulate 
from a broad, compressed base, terminal and 
ascending; stigma 2.5-3 mm long; endocarp in 
lower Yd; stem near the apex 13 mm in diam- 
eter; leaf with the caudate tip 1.3 mm wide, the 
marginal prickles near the base 3-6 mm apart, 
the nearby midrib below with prickles 0.5-1 
mm long, and at midsection the midrib un- 
armed. P. assamensis has the style 5-6 mm long, 
slender subulate, the apical drupes with terminal 
styles, the middle and lower ones with lateral 
styles; stigma 4-5 mm long; endocarp in lower 
V% stem near the apex 6-7 mm in diameter; 
leaf with the caudate tip 0. 5-0.7 mm wide, the 
marginal prickles near the base 14-26 mm apart, 
the nearby midrib below with prickles 2-2.5 mm 
long, and at midsection the midrib below with 
prickles 0. 7-0.8 mm long. 
SPECIES FROM THAILAND 
Section Aero stigma 
Pandanus retroaculeatus sp. nov. (sect. Acro- 
stigma ) . 
Fig. 226 
DIAGNOSIS HOLOTYPI: Planta 2 m alta, foliis 
circa 64 cm longis 2 cm latis supra viridibus 
infra pallide viridibus chartaceis excepta in basi 
subcoriacea supra midnervum sulcatis 2-plicatis 
ligulatis sed in dm ultimo in apice subulato 
diminuentibus (apex deest) in sectione me- 
dial! cum 28 nervis parallelis secundariis in quo- 
que latere ubique prominentibus ad apicem 
nervis tertiis obliquis remotis evidentis basi am- 
plexicauli et inermi conspicue nervosa ex 6 cm 
marginibus cum aculeis 1.5-2 mm longis 2-5 
mm separatis subulatis luteis inferis divergenti- 
bus, midnervo infra ex 9 cm cum aculeis 3.5-5 
mm longis 5-12 mm separatis crassiter subulatis 
valde reflexis basi incrassata, in sectione mediali 
marginibus cum aculeis 0.8-1 mm longis 2-5 
mm separatis subulatis adscendentibus, mid- 
nervo angusto et elevato cum aculeis 0. 5-0.8 mm 
longis 3-5 mm separatis subulatis adscendenti- 
bus, proxima apicem marginibus cum serrulis 
0.2-0.4 mm longis 0.5-1. 5 mm separatis, mid- 
nervo infra cum serrulis 0.6-0.8 mm longis 2-3 
mm separatis, in pagina supera ex medio ad 
apicem plicis binis cum aculeis 0.3-0. 5 mm 
longis 1-3 mm separatis subulatis adscendenti- 
bus, inflorescentia foeminea erecta folioso-brac- 
teata cum syncarpio solitario, pedunculo 13 cm 
longo 3-4 mm diametro 3-laterato, bracteis 
floraceis circa 9 luteis inferiore 7.5 cm longa 1 
cm lata lineari-lanceolata marginibus mediae 
ultimae cum aculeis pallidis congestis 0. 1-0.2 
mm longis, syncarpio (immaturo) 2 cm longo 
18 mm diametro ovoideo cum circa 288 drupis 
eis cum pileo 6-8 mm longo basi 3 mm alta 
5-angulosa lanceoloidea laevi et cum sulca pro- 
Fig. 225. Pandanus assamensis St. John, from holotype. a, fruiting branch, lateral view, X 1 ^ drupe, 
lateral view, X 1; c, drupe, longitudinal median section, XI \ d, drupe, lateral view, X 4; e, drupe, longi- 
tudinal median section, X 4; /, drupe, style, and stigma, apical view, X 4; g, leaf base, lower side, XI h, leaf 
middle, lower side, X 1 \ i, leaf apex, lower side, X 1- 
