526 
fundo ex stigmate descendant!, stylo 3-5 mm 
longo subnlato subarcuato vel recto osseoso sub- 
luteo, stigmate 3.5-5 mm longo linear! distal! 
obscure brunneo papillose, corpore 2.5-3 mm 
longo immature. 
DIAGNOSIS OF HOLOTYPE: Plant 2 m tall; 
leaves about 64 cm long, 2 cm wide, green 
above, pale green below, towards the base sub- 
coriaceous, the rest chartaceous, sukate above 
the midrib, 2 -pleated, ligulate but in the last 
dm tapering to a subulate apex ( the tip lacking 
in ours), at midsection with 28 parallel second- 
ary veins, these prominent throughout, towards 
the apex the remote oblique tertiary cross veins 
visible, the base amplexicaul and unarmed, 
prominently veiny, beginning at 6 cm the mar- 
gins with prickles 1.5-2 mm long, 2-5 mm 
apart, subulate, yellowish, the lowest ones diver- 
gent, the midrib below beginning at 9 cm with 
prickles 3.5-5 mm long, 5-12 mm apart, stout 
subulate, sharply reflexed, the base thickened; at 
midsection the margins with prickles 0.8-1 mm 
long, 2-5 mm apart, subulate, ascending; the 
midrib below narrow and raised, with prickles 
0. 5-0.8 mm long, 3-5 mm apart, subulate, as- 
cending; near the tip the margins with serf illa- 
tions 0.2-0.4 mm long, 0.5-1. 5 mm apart; the 
midrib below with serrulations 0.6-0.8 mm 
long, 2-3 mm apart; on the upper surface from 
the middle to the tip the pleats with prickles 
0.3-0. 5 mm long, 1-3 mm apart, subulate, 
ascending; pistillate inflorescence erect, leafy 
bracted, bearing one syncarp; peduncle 13 cm 
long, 3-4 mm in diameter, 3 -sided; floral bracts 
about 9, yellow, the lowest 7.5 cm long, 1 cm 
wide, linear-lanceolate, the margins of the outer 
half with pale, crowded prickles 0. 1-0.2 mm 
long; syncarp (young) 2 cm long, 18 mm in 
diameter, ovoid, bearing about 288 drupes, these 
with pileus 6-8 mm long, its base 3 mm high, 
5 -angled, lanceoloid, smooth, with a deep fur- 
row continuing down from the stigma; style 3-5 
mm long, subulate, slightly arcuate or straight, 
bony, yellowish; stigma 3.5-5 mm long, linear, 
distal, dark brown, papillose; body 2.5-3 mm 
PACIFIC SCIENCE, Vol. XIX, October 1965 
long, too immature to reveal its structure. 
HGLOTYPUS: Thailand, Ronpibun Hill, Na- 
kawn Srithamrat, jungle, 2,000 ft alt, Eryl Smith 
485 (BK). 
DISCUSSION : P. retroaculeatus is a member of 
the section Aero stigma, as is its closest relative, 
P. Toinu St. John, also of Thailand, a species 
with the pileus 5 mm long; stigma 2-2.5 mm 
long; stems 15 cm tall; leaves 14 mm wide, firm 
chartaceous, at midsection with 19-20 parallel 
secondary veins in each half, the tertiary cross 
veins visible nearly throughout, beginning at 5 
cm from the base the margins with prickles 
1- 1.5 mm long, 2-5 mm apart, lance-deltoid, 
and flat. P. retroaculeatus has the pileus 6-8 mm 
long; stigma 3.5-5 mm long; stems 1.5 m tall; 
leaves 20 mm wide, chartaceous except for 
the subcoriaceous base, at midsection with 28 
parallel secondary veins in each half, the tertiary 
cross veins oblique, remote, visible towards the 
apex, beginning at 6 cm from the base the mar- 
gins with prickles 1.5-2 mm long, 2-5 mm 
apart, and stout. 
The new epithet is from the Latin retrorsus, 
backward, and aculeatus, with prickles, and it 
is given with allusion to the strong, retrorse 
prickles of the midrib. 
Panddmis Toinu sp. nov. (sect. Act o stigma ) . 
Fig. 227 
NOM. VEKN.: "t6i nu.” 
DIAGNOSIS HOLOTYPI: Planta erecta aggre- 
gata, caii lib us 15 cm altis 8 mm diametro, foliis 
72 cm longis 14 mm lads fume chartaceis su- 
pra viridibus infra pallide viridibus 1-sulcatis 2- 
plicatis in sectione AA-formatis ligulatis in apice 
in 8 cm ultimis sensim diminuentibus in apice 
subulate 1 cm longo in sectione medial! cum 
19-20 nervis parallels secundariis in quaque 
media ei§ ubique evidentis et reticulis oblongis 
elongatis formantibus in media ultima plicis su- 
pra cum subulato-serrulis 0. 5-0.9 mm longis 
2- 4 mm separatis basi amplexicauli inermi sed 
nervis longitudinalibus prominentissimis ex 5 
Fig. 226. Pandanus retroaculeatus St. John, from holotype. a, fruiting branch, lateral view, X I; b, c, drupe, 
lateral view, X 1; d, drupe, lateral view X 10; e, drupe, longitudinal median section, X 10; f, g, pileus, style, 
and stigma, lateral view, 3^ 10; h, leaf base, lower side, .X 1; h leaf middle, lower side, X 1; j, leaf apex, 
upper side, X 1; leaf apex, lower side, X 1. 
