528 
PACIFIC SCIENCE, Vol. XIX, October 1965 
cm marginibus cum aculeis 1-1.5 mm longis 
2-5 mm separatis lanceo-deltoideis complanatis 
luteis adscendentibus, midnervo infra ex 5.5 cm 
cum aculeis 0.7-1. 5 mm longis 2-8 mm sepa- 
ratis subulato-conicis infimis erectis alteris re- 
flexis, in sectione mediali marginibus cum 
subulato-serrulis 0.7-0. 9 mm longis 2-3 mm 
separatis, midnervo infra cum serrulis 0.5 mm 
longis 4-8 mm separatis vel nullis, ad apicem 
marginibus et midnervo infra cum serrulis 0.1 
mm longis 1-2 mm separatis, inflorescentia foe- 
mi nea erecta foliosa cum syncarpio solitario, 
pedunculo 8 cm longo 4 mm diametro trigono, 
syncarpio (immaturo) 2 cm longo 18 mm 
diametro ovoideo-subgloboso cum circa 240 
dr up is eis immaturis 7-8 mm longis, pileo 5 
mm longo obliquiter lanceoloideo proxime cur- 
vato osseoso luteo basi 2 mm alta 1.7—2 mm lata 
obliquiter pyramidali prominente 5-angulosa 
glauca, stylo 3 mm longo crassiter subulato 
arcuato lucido subtereto, stigmate 2-2.5 mm 
longo late lineari cinnamomum-brunneo papil- 
lose subterminali, corpore 2.5-3 mm longo ob- 
lanceoloideo immaturo contracto. 
DIAGNOSIS OF HOLOTYPE: Plant erect, form- 
ing tufts; stems about 15 cm tall, 8 mm in 
diameter; leaves 72 cm long, 14 mm wide, firm, 
chartaceous, green above, pale green below, with 
one central furrow, 2 -pleated, in section M- 
shaped, ligulate, at tip gradually narrowed in 
the last 8 cm to a 1 cm subulate apex, at mid- 
section with 19-20 parallel secondary veins in 
each half, these prominent throughout, the ter- 
tiary cross veins visible nearly throughout, form- 
ing a network with the meshes elongate oblong, 
on the outer half the pleats on the upper sur- 
face with subulate-serrulations 0. 5-0.9 mm long, 
2-4 mm apart, the base amplexicaul, unarmed, 
but the longitudinal veins very prominent, be- 
ginning at 5 cm the margins with prickles 1—1.5 
mm long, 2-5 mm apart, lance-deltoid, flat, yel- 
low, ascending; the midrib below beginning at 
5.5 cm with prickles 0.7-1. 5 mm long, 2-8 mm 
apart, subulate conic, the lowest erect, the others 
becoming reflexed; at midsection the margins 
with subulate-serrulations 0.7-0.9 mm long, 2-3 
mm apart; the midrib below with serrulations 
0.5 mm long, 4-8 mm apart, or none; near the 
tip the margins and midrib below with serru- 
lations 0.1 mm long, 1-2 mm apart; pistillate 
inflorescence erect, leafy, bearing a single syn- 
carp; peduncle 8 cm long, 4 mm in diameter, 
trigonous; syncarp (immature) 2 cm long, 
18 mm in diameter, ovoid-subglobose, bearing 
about 240 drupes, these (immature and the body 
little developed) 7-8 mm long; pileus 5 mm 
long, obliquely lanceoloid, proximally curved, 
bony, yellow, the base 2 mm high, L7-2 mm 
wide, obliquely pyramidal, prominently 5 -an- 
gled, glaucous; style 3 mm long, thick subulate, 
arcuate, shining, subterete; stigma 2-2.5 mm 
long, broad linear, cinnamon-brown, papillose, 
reaching quite to the tip; body 2.5-3 mm long, 
oblanceoloid, shrunken, immature. 
HOLOTYPUS: Thailand, Padang Besar, 50 m 
alt, light evergreen forest, Dec. 25, 1927, A. F. 
G. Kerr 13,639 (bk). 
DISCUSSION: P. T oinu is a member of the 
section Acrostigma, as is its closest relative, P. 
similis Craib, of Thailand, a species with the 
pileus 5 mm wide, the base hemispheric; style 
flat; leaves 1 m long, 14—19 mm wide, near the 
base the midrib with prickles 1.5-2 mm long, 
15-20 mm apart, reflexed, at midsection the 
margins with white serrations 0.4-0. 5 mm long, 
and 3-6 mm apart. P. Toinu has the pileus 1.7- 
2 mm wide, the base obliquely pyramidal; style 
subterete; leaves 72 cm long, 14 mm wide, near 
the base the midrib with prickles 0.7-1. 5 mm 
long, 2-8 mm apart; at midsection the margins 
with yellow subulate-serrulations 0.7-0. 9 mm 
long, and 2-3 mm apart. 
The specific epithet is the vernacular name 
of the species, used as a noun in apposition. 
Section Rykia 
Pandanus acaulescens sp. nov. (sect. Rykia). 
Fig. 228 
Fig. 227 . Pandanus Toinu St. John, from holotype. a, fruiting branch, lateral view, XI b, c, drupe, 
lateral view, X 1; ^ drupe, lateral view, X 10; e, drupe, longitudinal median section, X 10; f, g, style, and 
stigma, lateral view, X 10; h, leaf base, lower side, X 1; -i, leaf middle, lower side, X 1; j, leaf apex, upper 
side, X 1; k, leaf apex, lower side, X 1- 
