530 
PACIFIC SCIENCE, Vol. XIX, October 1965 
DIAGNOSIS HOLOTYPI: Planta acaulescens a g- 
gregata, foliis 2.54-3.5 m longis circa mediam 
6.1 cm latis proxima basem 4 cm latis subtiliter 
coriaceis supra midnervum late sulcatis 2-plica- 
tis in sectione mediali cum 49 nervis parallelis 
secundariis in quaque media nervis tertiis in 
media ultima conspicuis et reticulis quadratis 
formantibus lamina ligulata in apice trigono 
subulate 15 cm longo diminuentibus eo in 
puncto 10 cm ex apice 1.5 mm lato basi am- 
plexicauli inermi crassa lucida rubro-brunnea, 
marginibus ex 16 cm cum aculeis 1.5-2. 5 mm 
longis 5-10 mm separatis subulatis pallidis ad- 
scendentibus, midnervo infra ex 14 cm cum 
aculeis 2-3 mm longis 11-18 mm separatis 
arcuatis subulatis recurvatis per 16 cm extentis 
deinde ad Va ultima inermi, in sectione mediali 
marginibus cum serrulis 0.7-1 mm longis 4-11 
mm separatis, in apice subulato marginibus et 
midnervo infra cum serrulis 0.6-1 mm longis 
1-4 mm separatis, inflorescentia foeminea erecta 
cum syncarpio solitario, pedunculo 56 cm longo 
trigono subclavato in apice 15 mm diametro 
media ultima folioso-bracteato, bracteis syncar- 
pium includentibus interioribus marcescentibus 
et in stigmatibus affixis, bracteis superis 17 cm 
longis 6 cm latis marginibus superis et mid- 
nervo infra serrulatis, syncarpio 14.5 cm longis 
9.3 cm diametro ellipsoideo 3-laterato cum circa 
696 drupis in lineis verticalibus, dr up is 32-33 
mm longis 7-10 mm latis 6-10 mm crassis ob- 
scure viridibus crassiter cuneatis 6-angulosis 
parte V3 supera libera corpore 25-26 mm longo 
lateribus planis, pileo cum basi 4-5 mm alta 
subrotundata-pyramidali exlucida, stylo 4-6 mm 
longo longiter cornu formi osseoso mahogani- 
brunneo lucido infra tereto supra complanato 
valde proxime curvato plerumque V 5 -V 5 bifido 
ramis plerumque divergentibus (stylo druparum 
terminalium subulato integro), stigmate 3-3.5 
mm longo lanceolato proximo brunneo papil- 
loso, endocarpio mediali elliptico-oblanceoloideo 
osseoso brunneo lateribus lateralibus 1 mm cras- 
sis intra laevi et lucido apice in projectione 
lanceo-subulato extenso, semine 13 mm longo 
oblanceo-ellipsoideo, mesocarpio apicali caver- 
noso cum membranis medullosis transversis al- 
bis, mesocarpio basali fibroso et carnoso. 
DIAGNOSIS OF HOLOTYPE: Acaulescent, tufted 
plant; leaves 2.54-3.5 m long, 6.1 cm wide a 
little beyond the middle, near the base 4 cm 
wide, thin coriaceous, broadly furrowed above 
the midrib, 2 -pleated, at midsection with 49 
parallel secondary veins in each half, the tertiary 
cross veins conspicuous in the outer half, mak- 
ing a reticulum of squarish meshes, blade ligu- 
late, tapering to a 15 cm trigonous, subulate 
tip, this 10 cm down 1.5 mm wide, the base 
amplexicaul, unarmed, thick, shining, reddish 
brown, the margins beginning at 16 cm with 
prickles 1.5-2. 5 mm long, 5-10 mm apart, sub- 
ulate, pale, ascending; the nearby midrib below 
beginning at 14 cm with prickles 2-3 mm long, 
11-18 mm apart, arcuate, subulate, recurved, 
these extending for only 16 cm, then the midrib 
unarmed till the last quarter; at midsection the 
margins with serrulations 0.7-1 mm long, 4-11 
mm apart, on the subulate apex the margins and 
midrib below with serrulations 0.6-1 mm long, 
1-4 mm, -apart; pistillate inflorescence erect, with 
1 syncarp; peduncle 56 cm long, trigonous, sub- 
clavate, at apex 15 mm in diameter, the upper 
half leafy bracted, the bracts enclosing the syn- 
carp and the inner ones becoming marcescent 
and impaled on the sharp stigmas; upper bracts 
17 cm long, 6 cm wide, their upper margins and 
midrib serrulate; syncarp 14.5 cm long, 9.3 cm 
in diameter, ellipsoid, 3 -sided, bearing in ver- 
tical rows about 696 drupes, these dark green, 
32-33 mm long, 7-10 mm wide, 6-10 mm 
thick, cuneate, 6-angled, upper l A free, the body 
25-2 6 mm long, its sides flat; pileus with the 
base 4-5 mm high, slightly rounded pyrami- 
dal, dull; style 4-6 mm long, hornlike, bony, 
mahogany-brown, shining, terete below, flat- 
tened above, sharply proximally curved, mostly 
Fig. 228. Pandanus acaulescens St. John, from holotype. a, syncarp, lateral view, X Vl\ b, drupe, lateral 
view, X 1; c, drupe, longitudinal median section, X 1 \ d, i, j, k, h o, lateral drupes, apical view, X 1; e, 
f, g, h, apical drupes, apical view, X 1; Pi drupe, lateral view, X 4; q, drupe, longitudinal median section, 
X 4; r, style and stigma of lateral drupe, lateral view, X 4; s, style and stigma of apical drupe, lateral view, 
X 4; t, leaf base, lower side, XI \ u, leaf middle, lower side, X 1; ^ secondary and tertiary venation of leaf 
middle, lower side, X 4; w, leaf apex, lower side, X 1- 
