) 
532 
bifid Vi- 2 /^ way, the forks mostly divergent 
(the apical drupes with the style simple, subu- 
late); stigma 3-3.5 mm long, lanceolate, on the 
proximal face of the lobes, united below, brown, 
papillose; endocarp median, elliptic-oblanceo- 
loid, bony, brown, the lateral walls 1 mm thick, 
the inner surface smooth, shining, the apex pro- 
longed upwards into a lance-subulate projec- 
tion; seed 13 mm long, oblance-ellipsoid; apical 
mesocarp cavernous, with transverse, white, 
medullary membranes; basal mesocarp fibrous 
and fleshy. 
HOLOTYPUS: Thailand, Chiengmai, Tat Noi, 
common in dry waterway, 900 m alt, tufted 
plant, no stems, 19 April I960, Tem Smitinand 
6,746 (bkf). 
DISCUSSION: P. acaulescens is a member of 
the section Rykia, as is its closest relative, P. 
penetrans St. John, a species found in the same 
general locality, and one which has the infruc- 
tescence pendent; syncarp 12 cm in diameter; 
drupes 33-35 mm long, 9—1 1 mm wide, upper 
l A free; pileus 12-15 mm high, the base ob- 
lance-pyramidal; style 7-8 mm long; endocarp 
submedian; leaves 5-5.6 cm wide near the base, 
at midsection with 43 parallel secondary veins 
in each half, at midsection the margins with 
prickles 1.5-2. 5 mm long, 13-22 mm apart, 
subulate, the nearby midrib below with prickles 
3.5-5 mm long, 38-70 mm apart, stout subu- 
late, reflexed; tree up to 9 m tall, 8 cm in 
diameter, the bark with ascending adventitious 
rootlets. P. acaulescens has the infructescence 
erect; syncarp 9.3 cm in diameter; drupes 32-33 
mm long, 7—10 mm wide, upper VS free; pileus 
8-1 1 mm high, the base slightly rounded pyram- 
idal; style 4-6 mm long; endocarp median; 
leaves 4 cm wide near the base, at midsection 
with 49 parallel secondary veins in each half, at 
midsection the margins with serrulations 0.7-1 
mm long, 4-11 mm apart, the nearby midrib 
PACIFIC SCIENCE, Vol. XIX, October 1965 
below unarmed; and the plant acaulescent. 
The new epithet is the Latin acaulescens, 
without a stem, and is here given as a descrip- 
tive name. 
Pandanus ligulatus sp. nov. (sect. Rykia). 
Fig. 229 
DIAGNOSIS HOLOTYPI: Truncus 3 m altus, 
foliis 3.95 m et plus longis 4 cm latis subcori- 
aceis apparente pallide viridibus supra mid- 
nervum late sulcatis 2-plicatis ligulatis apice 
perdito, in sectione mediali cum 46 nervis paral- 
lels secundariis in quoque dimidio ad apicem 
praecipue infra nervis tertiis conspicuis et 
oblongos angustos formantibus basi arnplexi- 
cauli et inermi cupraceo sed ex 13-18 cm mar- 
ginibus cum spinis 4-4.5 mm longis 25-45 mm 
separatis crasse arcuato-subulatis adscendentibus 
brunneis, midnervo infra cum spinis simulanti- 
bus sed crassioribus et reflexis, in sectione medi- 
ali marginibus cum aculeis 2-2.5 mm longis 
15-21 mm separatis subulatis adpresse adscen- 
dentibus apicibus brunneis, midnervo infra cum 
aculeis simulantibus adscendentibus sed 30-33 
mm separatis, in apice marginibus cum subulato- 
serrulis 1—1.5 mm longis 3-5 mm separatis, 
midnervo infra cum serrulis simulantibus 8-12 
mm separatis, infructescentia pendenti cum syn- 
carpio solitario, pedunculo 70 cm longo 1.3 
cm diametro trigono folioso-bracteato, bracteis 
superis 20 cm longis 5 cm latis foliaceis mar- 
ginibus cum subulato-serrulis 0.5 mm longis 
3-5 mm separatis, syncarpio 11 cm longo 7.5 cm 
diametro late ellipsoideo viridi, drupis 27-32 
mm longis 6-8 mm latis 5-7 mm crassis ob- 
longo-oblanceoloideis eis lateralibus et subba- 
salibus minime compressis (illis apicalibus parve 
majoribus et paucis cum stylo subulato integri 
arcuato), pileo 10-13 mm longo basi 4-7 mm 
alto anguste pyramidali-conico 5-6-anguloso 
minute papilloso, stylo 6-7 mm longo osseoso 
brunneo lucido supra mediam compresso prox- 
FlG. 229- Pandanus ligulatus St. John, from holotype. a, drupe, lateral view, X 1 ^ drupe, longitudinal 
median section, X 1; c, drupe apex, apical view, X 1; d, e, f, g, h, style and stigma of lateral drupes, lateral 
view, X 1; i, j, style and stigma of apical drupes, lateral view, X 1; k, drupe, lateral view, X 4; l, drupe, 
longitudinal median section, X 4; m, style, distal view, X 4; n, style and stigma, proximal view, X 4; o, style, 
distal view, X 4; p, leaf base, lower side, X 1; leaf middle, lower side, XI \ r, secondary and tertiary vena- 
tion at middle, lower side, X 4; s, leaf apex, lower side, X 1- 
