536 
PACIFIC SCIENCE, VoL XIX, October 1965 
longis adventivis paucis spiniformatis adpresse 
adscendentibus, radicibus fulturosis nullis vel 
in truncis prostratis 3-5 dm longis 1 cm diame- 
tro paucis laevibus vel parve spinosis, foliis 
2.73-2.9 m longis 6-6.3 cm latis in media sed 
proxima basem 5-5.6 cm latis subcoriaceis supra 
obscure olivaceo-viridibus infra pallide viridi- 
bus in basi clausa cupracea supra midnervo late 
1-sulcatis 2-plicatis in sectione mediali cum 43 
nervis parallelis secundariis in quoque medio 
in dimidia ultima nervis tertiis conspicuis in 
lateribus ambis laminis ligulatis sed in basi sub- 
cuneatis et in apice subulato trigono gracili 
breviter diminuentibus eo 10 cm ex apice 1 mm 
lato basi amplexicauli et inermi sed marginibus 
ex 10-11 cm cum aculeis 3.5-4 mm longis 11- 
18 mm separatis subulatis validibus subadscend- 
entibus subviridibus vel albis apicibus brunneis, 
midnervo infra ex 18 cm cum aculeis 4-5 mm 
longis - 15-25 mm separatis crasse subulatis sub- 
reflexis in sectione mediali marginibus cum 
aculeis 1.5-2. 5 mm longis 13-22 mm separatis 
subulatis adpresse adscendentibus, midnervo in- 
fra cum aculeis 3.5-5 mm longis 38-70 mm 
separatis crasse subulatis reflexis, in apice subu- 
lato marginibus et midnervo infra cum serrulis 
0. 3-0.5 mm longis 1-6 mm separatis, inflores- 
centia foeminea terminali pendenti cum syncar- 
pio solitario, pedunculo 65 cm longo subclavato 
in apice 2 cm diametro trigono folioso-bracteato 
bracteis superis 80 cm longis 7 cm latis, bractea 
supera 17 cm longa 5.5 cm lata marginibus 
superis serrulatis, syncarpio cum bracteis marce- 
scentibus subclausis, syncarpio 14 cm longo 12 
cm diametro late ellipsoideo valde 3-laterato 
cum circa 576 drupis eis 33-35 mm longis 9-11 
mm latis 7-10 mm crassis oblongo-lanceoloideis 
5-7-angulosis (drupis terrninalibus parve ma- 
joribus et cum stylo subulato arcuato integro) 
eis lateralibus viridibus subglaucis lateribus 
planis laevibus parte l A supera libera, pileo 12- 
15 mm alto cum basi 7-10 mm alta 5-7-angu- 
losa oblanceo-pyramidali lateribus laevibus, stylo 
7-8 mm longo cornoso obscure brunneo lucido 
compresso adscendenti turn valde proxime cur- 
vato l A- 2 A furcato furcis spinosiformibus ple- 
rumque divergentibus et subaequalibus, stigmate 
4-5 mm longo lanceolato bifurcato brunneo 
papilloso paene ad apices extento, drupis apicali- 
bus cum stylo 5-6 mm longo subulato erecto 
recto vel subarcuato, endocarpio submediali cor- 
pore 14 mm longo osseoso brunneo in apice cum 
extensioni subulato 8 mm longo lateribus later- 
alibus 1-1.5 mm crassis intra lucidis, semine 13 
mm longo ellipsoideo, mesocarpio apicali caver- 
noso cum membranis albis medullosis multis, 
mesocarpio basali parvo fibroso et carnoso. 
DIAGNOSIS OF HOLOTYPE: Tree, up to 9 m 
tall, 8 cm in diameter; bark brown, with a few, 
well spaced, adventive rootlets, up to 1 cm long, 
appressed ascending, almost spine-like; prop 
roots none on erect trunks, but on fallen trunks 
rare and 3-5 dm long, 1 cm in diameter, smooth 
or with a few remote, small spines; leaves 2.73- 
2.9 m long, 6-6.3 cm wide at the middle, 5-5.6 
cm wide near the base, subcoriaceous, above 
dark olive green, below pale green, the covered 
auriculate base copper-colored, broadly furrowed 
above the midrib, 2 -pleated, at midsection with 
43 parallel secondary veins in each half, in the 
outer half the tertiary cross veins conspicuous 
on both sides, a little sinuous, making a pat- 
tern of squares or short oblongs, blade ligulate, 
though slightly tapering to the base and short 
tapering to the 10-15 cm, trigonous, slender 
subulate apex, this 10 cm down 1 mm wide, the 
base amplexicaul and unarmed, but the margins 
beginning at 10-11 cm with prickles 3.5-4 mm 
long, 11-18 mm apart, stout subulate, slightly 
ascending, greenish to whitish and brown- 
tipped; the midrib below beginning at 18 cm 
with prickles 4-5 mm long, 15-25 mm apart, 
stout subulate, slightly reflexed; at midsection 
the margins with prickles 1.5-2. 5 mm long, 13- 
22 mm apart, subulate, appressed ascending; the 
midrib below with prickles 3.5-5 mm long, 38- 
70 mm apart, stout subulate, reflexed; on the 
subulate apex the margins and midrib below 
with serrulations 0.3-0. 5 mm long, 1-6 mm 
apart; pistillate inflorescence terminal, pendent, 
bearing one syncarp; peduncle 65 cm long, 
slightly clavate, at apex 2 cm in diameter, trigo- 
nous, leafy bracted; upper bracts 80 cm long, 
7 cm wide, the innermost 17 cm long, 5.5 cm 
Fig. 231. Pandanus penetrans St. John, from holotype, syncarp, lateral view, X 1- 
