Bathyal Amphipoda— -BARNARD 
327 
the second; head with accessory lobe at antero- 
lateral corner behind which is the sharp tooth 
generally present in Melitas; eyes present, with 
well-developed ommatidea but bleached to ochre 
in alcohol; article 1 of mandibular palp not pro- 
duced; inner lobes of lower lip well developed; 
coxa 1 not produced forward; articles 5 and 6 
of gnathopod 1 equal in length, finger normal; 
palm of male gnathopod 2 oblique, with three 
processes, finger overriding palmar edge and 
slightly longer than palm, article 4 with distal 
tooth; pereopods 3-5 rather short and stout, 
posterodistal comers of article 2 not strongly 
produced; pleonal epimera rounded or quadrate 
at posterior lower comers; pleonal segments 1-5 
increasingly strongly dentate dorsally, on seg- 
ment 1 there are two obsolescent teeth, on 
segment 2 there are three obtuse teeth, on seg- 
ment 3 there are five teeth, on segment 4 there 
are three teeth, the middle-most of which is 
more erect than the two laterals, on segment 5 
there are three teeth, the middle of which is 
shortest but possibly because of damage; hence 
the pleonal tooth formula is (2 ) -3-5-3— 3-0. 
HOLOTYPE: USNM no. 108631, male, 12 mm. 
Unique. 
TYPE LOCALITY: Chogafka Cove, Kodiak Is- 
land, Alaska, July 10, 1880, W. H. Dali, no. 
3371 (1948). 
RELATIONSHIP: This species is related to M. 
valida Shoemaker (1955) but differs by the 
quadrate third pleonal epimeron, the smaller 
forward extension of the first coxa, and the 
larger teeth of the urosome. It differs from M. 
dentata (Kroyer) (see Sars, 1895: pi. 181) by 
the quadrate third pleonal epimeron, the un- 
produced first mandibular palp article, and the 
accessory lobe at the lower anterior corner of 
the head. From M, gladiosa Bate ( Chevreux and 
Fage, 1925:233) it differs by the unproduced 
first to third pleonal epimera and the male 
second gnathopodal palmar armature. From M. 
pallida Sars (1886: pi. 15) the new species 
differs by the unproduced second and third 
pleonal epimera and the shape of the head. 
FAMILY PHOTIDAE 
GENUS Photis Kroyer 
Photis chiconola, new species 
Fig. 9 
DIAGNOSIS OF MALE: Coxae 1 and 2 similar 
to coxae 3 and 4, longer than broad; third coxa 
not wider than second; gnathopod 1 with palm 
slightly excavate, distinctly defined by a shallow 
bump armed with a spine; palm of gnathopod 2 
oblique, with a deep, narrow conical invagina- 
tion, the defining tooth slender but not reaching 
a transverse line from the dactylar hinge, the 
palmar process near finger hinge not protruding 
strongly, article 7 slender, tapering evenly, with 
several inner serrations and articulated spines, 
the largest occurring distally and forming a 
slight notch, tip of article 7 strongly overlapping 
defining tooth; article 2 of gnathopod 2 poorly 
produced anterodistally, its lateral face with a 
single, obscure stridulating ridge. 
FEMALE: Palm of gnathopod 2 slightly ex- 
cavate, its article 7 overlapping palm, antero- 
distal end of article 2 with a small, narrow 
process. 
HOLOTYPE: USNM no. 108632, male, 5 mm. 
TYPE LOCALITY: Albatross Station 4530, 
Monterey Bay, California, Pt. Pinos Lighthouse, 
S 78° E, 6.8 miles, 847-755 fms, beam trawl 
on soft gray mud, May 27, 1904. 
MATERIAL : Station 4530 (25 specimens). 
relationship: This species is most closely 
related to Photis conchicola Alderman (see J. L. 
Barnard, 1962a) and the differences herein 
noted may be those of only racial value. Photis 
chiconola differs from P. conchicola especially 
by the slender dactyl of the female gnathopod 2, 
which strongly overlaps the palm, in contrast to 
the short, stout dactyl fitting the palm in P. 
conchicola. Articles 3-6 of pereopods 3-5 are 
more slender in P. chiconola, the dactyl of male 
gnathopod 1 is longer, and the male coxa 2 is 
longer than broad but this probably is a feature 
of its juvenility. The distal lobe of article 2 on 
the female second gnathopod is small and slender 
in contrast to that of female P. conchicola. 
The new species is related to P. reinhardi 
Kroyer (see Sars, 1895 : pi. 202), differing from 
it by the narrower palmar excavation of male 
gnathopod 2, which may be a feature of juvenil- 
ity, but the dactyl strongly overlaps the palm, 
indicating sufficient difference. 
