362 
PACIFIC SCIENCE, Vol. XVIII, October 1964 
including inferior angle; upper surface of sec- 
ond and third teeth supporting a few widely 
spaced spinules; fourth tooth with a few small 
accessory spines near base; superior margin of 
mandible with row of numerous paired setae; 
inferior margin supporting approximately eight 
spines; a group of several spines near superior 
angle, and a lower group of numerous short 
spines in groups or singly similar in arrange- 
ment to that seen in Heteralepas gigas (An- 
nandale, 1905, pL 8, fig. 3b). First maxilla 
(Fig. 3 E) with three major spines above and 
approximately seven major spines below promi- 
nent notch; a few long spines in a group along 
superior margin, and two long spines near the 
base of first major spine; cutting edge composed 
of notch and superior spines equal in length to 
cutting edge below notch; surface clothed with 
numerous slender spinules arranged in groups 
and rows; notch supporting a few strong spines; 
inferior margin with approximately eight long 
spines. Second maxilla (Fig. 3 F) broadly 
rounded; superior margin with minute and 
widely spaced serrations and a few spines; inner 
margin supporting long plumose spines in two 
groups, one at the superior and the other at the 
inferior angle; space between these two groups 
without spines (a condition occasionally seen 
in Heteralepas japonica according to Nilsson- 
Cantell, 1927). At least two very long pinnate 
spines on swelling at proximal portion of in- 
ferior margin. 
Insertion of cirrus I widely separated from 
cirrus II, attached to posterior margin of pro- 
soma; cirrus I approximately one-half length of 
cirrus II, supporting a single posteriorly directed 
ensiform filamentary appendage at its base; cirri 
II-VI essentially equal in length; inner ramus of 
cirri V and VI atrophied; pedicles of all cirri 
faintly squamous; long setae of all cirri finely 
pinnate, but becoming progressively less so 
posteriorly; inner ramus of cirrus I somewhat 
shorter than the outer, and more heavily clothed 
with setae; rami equal in width, arrangement of 
.setae lasiopod (as many as 12 long, strong setae 
in staggered transverse whorl), with at least 
one major spine at each articulation along the 
greater curvature on intermediate articles; outer 
ramus of cirrus II slightly longer than the inner, 
setae arranged normally (cf Fig. 3 G)\ setal 
arrangement of inner ramus intermediate be- 
tween normal form and that of outer ramus of 
cirrus I; inner rami of cirri V and VI atrophied, 
approximately one-fifth length of outer rami; 
proximal articles of all cirri tending to fuse; 
unfused proximal articles two-thirds wider than 
long, intermediate articles as wide as long, distal 
articles essentially one-half as wide as long; 
counts for cirri of the holotype UCMP 37863 
and a paratype (in parentheses) are given be- 
low. Unfortunately, the tips of many rami had 
been nipped off and no count could be made. 
Caudal appendages of five articles nearly as 
long as the first article of the pedicle of cirrus 
VI, last two articles supporting a few long 
spinules, apex with a tuft of several short and 
two long spinules. Penis (Fig. 3 H) weakly annu- 
lated, relatively short, slender, tapering gradually 
throughout its length; clothed with numerous 
long, soft setae and occasional short spines; apex 
truncate, supporting a tuft of short stiff bristles. 
TYPE DESIGNATION: Holotype: UCMP 37863. 
Paratype: UCMP 37864. Additional specimen: 
U.S. National Museum, Washington, D.C, no. 
109782. 
DISCUSSION: Although the species described 
here cannot be assigned to any of the known 
forms of Heteralepas s. str., it is similar to 
several, some of which are of uncertain status. 
These are: 
1. Heteralepas japonica ( Aurivillius, 1894) 
2. H. indica (Gruvel, 1902) 
3. H. gigas (Annandale, 1905) 
4. H. cygnus Pilsbry, 1907 
5. H.lankesteri (Gruvel, 1901 ) 
6. H. quadrata (Aurivillius, 1894) 
Except for H. quadrata, these forms generally 
occur on the spines of sea urchins. They are 
characterized externally by an ovoid, slightly 
compressed, relatively smooth capitulum, and 
protuberant or slightly protuberant lips. Except 
Cirrus: I 
Right outer ramus : 16(19) 
Right inner ramus : 10(13) 
Left outer ramus : 16(19) 
Left inner ramus : 11(13) 
II 
III 
IV 
-(37) 
-(42) 
-(41) 
29(27) 
-(-) 
-(39) 
31(39) 
-(-) 
-(40) 
25(31) 
-(-) 
-(-) 
V 
VI 
CA. 
-(-) 
35 (-) 
-(5) 
6(11) 
8(8) 
-(45) 
-(-) 
5 (-) 
10(10) 
-GD 
