Thoracic Cirripedia from a Guyot — Z ullo and Newman 
363 
for H. lankesteri from the West Indies, they are 
reported from the Pacific and Indian oceans. 
Nilsson-Cantell (1927) discusses variability 
in certain external and internal characteristics 
ordinarily utilized in distinguishing members of 
the genus. In an extensive study of H. japonica, 
he concluded that H. indica is but a form of 
this species and that the status of H. nicobarcia 
Annandale, H. gigas, and H. cygnus is subject to 
re-examination in the light of his findings. 
Broch (1931), Nilsson-Cantell (1938), and 
Utinomi (1958) list H. indica as a synonym of 
H. japonica, but the status of the other species 
has not been altered. Whatever the status of 
these species, the form described here differs 
from them in a number of ways, in that H. 
japonica and its allies have a rather large orifice, 
occupying essentially one-half the occludent 
margin of the capitulum, and the mandible, at 
least of H. japonica, has pectinations on the 
underside of the first, second, and third teeth. 
The orifice of H. lankesteri is a little less than 
one-half the height of the capitulum, but the 
mandible is essentially that of H. japonica. Re- 
maining then is H . quadrata. It generally occurs 
on Pcmulirus in the Pacific. Despite its similar 
appearance, it can be separated from the species 
described here by the presence of minute rivet- 
like structures (figured by Utinomi, 1949) on 
the penis. 
The nearly naked Lepadomorpha present a 
difficult problem for the systematist since, being 
unarmored, they lack a number of distinctive 
features customarily utilized in separating gen- 
era and species. Turning to a comparative study 
of the appendages, Pilsbry (1907) was able to 
separate Heteralepas s. 1. from Alep as, and to 
split the genus into two distinct subgenera: 
Heteralepas s. str. and Paralepas . However, he 
retained the heterogeneous assemblage Alepadi- 
nae. Nilsson-Cantell (1921) furthered our un- 
derstanding of this group by placing Heteralepas 
s. 1. in a separate subfamily, the Heteralepadidae. 
Further work sustained Pilsbry’s subgeneric dis- 
tinctions, and, as a consequence of his sugges- 
tion, the subgenera were finally recognized as 
separate genera (Newman, I960). 
At higher taxonomic levels the group would 
appear to be in fairly good order. However, as 
has been discussed above, this is not true as far 
as many of the species are concerned. In fact, 
it seems fair to say that we simply do not know 
what morphological and zoogeographic data are 
really applicable in making judgments at the 
specific level, there being so little information 
to go on. Aside from the work of Nilsson- 
Cantell (1927), no critical studies have been 
made. Thus it is difficult to establish a new form 
with any degree of certainty, for extensive col- 
lections would be required and they are simply 
not available. Therefore much of the time spent 
in study of this form was devoted to comparing 
characters with the known range of variability. 
Even so, with the extreme latitude allowed by 
synonymy, the present form could not be as- 
signed to any known species. 
The new species has been named H. mystaco- 
phora ("mustache bearer”) for the numerous 
soft setae clothing the bullate portion of the 
labrum or upper lip. This condition is indeed 
unusual, and it is difficult to conceive what 
function it may serve. Yet the palps are provided 
with long setae or spines, perhaps stronger and 
certainly fewer than is usual in cirripeds, and 
they are in a position to be drawn through the 
labral setae. Perhaps in this way material en- 
tangled there could be brought to the mouth 
field. 
suborder VERRUCOMORPHA Pilsbry 
FAMILY VERRUCIDAE Darwin 
GENUS Verruca Schumacher, 1817 
SUBGENUS Verruca s. str. 
Verruca Schumacher, Pilsbry, 1916, U.S. Nat. 
Mus. Bull. 93, p. 23. 
Eu-Verruca Broch, 1931, Vidensk. Meddel. 
Dansk naturhist. Foren., Bd. 91, p. 45. 
Verruca ( Verruca ) scrippsae Zullo, sp. nov. 
Figs. 4A-K 
Dimensions of holotype UCMP 347 1 3 : height 
of shell, 2.5 mm; carinorostral diameter of base, 
6 mm; lateral diameter of base, 4 mm; carinoros- 
tral diameter of orifice, 2.5 mm. 
Shell (Figs. 4 A, B) depressed, boxlike, colored 
white in dried specimen with adherent particles 
of yellow-brown epidermis; fixed plates steep, 
approaching perpendicular; movable valves flat, 
parallel to base; base of fixed plates thin, not 
