Four New Species, a New Genus, and a New Suborder of Hawaiian Fishes^ 
William A. Gosline^ 
The description of new taxa in a ’'Hand- 
book of Hawaiian Fishes” recently submitted 
for publication was considered inadvisable. 
Certain of these are grouped together here, 
i. e., three new species of Gobiidae {Quisqui- 
lius aureoviridis, Q. limbatosquamis , and Hazeus 
unisquamis), a new genus and species of Cal- 
lionymidae {Pogonymus pogognathus), and a 
new suborder of Perciformes (Schindlerioidei) 
erected for the fishes of the genus Schindleria. 
Family GOBIIDAE, Genus Quisquilius 
The two new species of Quisquilius de- 
scribed below can be assigned to that genus 
with considerable assurance, as the type spe- 
cies, Q. eugenms, is another Hawaiian form of 
which numerous specimens are available. The 
characteristics listed in the following para- 
graph are held in common by the two new 
species and Q. eugenius. 
No hairlike or flaplike appendages on the 
head or shoulder girdle. No serrations or 
spines on the preopercle. Maxillary not pro- 
longed posteriorly. No V-shaped notch in the 
tip of the lower jaw. No teeth on the vomer. 
Teeth in both jaws in several rows, those of 
the outer and inner rows enlarged and spaced 
at intervals. One or two fanglike teeth in the 
outer row at each anterolateral angle of the 
lower jaw. Tongue usually with a slight to 
broad notch in the tip. Cleft of the mouth 
more or less oblique. Both nostrils in tubes, 
the anterior at least twice as far from the pos- 
terior as from the groove behind the upper 
lip. No pores on the head. The rows of 
* Contribution No. 112 of the Hawaii Marine 
Laboratory in cooperation with the University of 
Hawaii, Department of Zoology and Entomology. 
Manuscript received December 18, 1957. 
2 Department of Zoology and Entomology, Univer- 
sity of Hawaii. 
papillae (not shown in Fig. 2) are in very 
uniform alignment. There is one pair on the 
posterior portion of the interorbital and two 
rows of about 5 papillae each on the flesh 
covering the premaxillary pedicels. A series of 
about 5 large, widely spaced papillae borders 
the orbit below. A row or pair of rows runs 
backward from the eye just above the upper 
border of the opercle. Two horizontal rows 
on cheek, the upper sometimes extending all 
the way across but the lower ending on mid^ 
die of cheek. A verticle row down front of 
operculum with two branches running back 
or down and back from it. One or two rows 
along the lower border of the preopercle. One 
row of papillae above and two below enclose 
the corner of the mouth. Two rows on the 
gular membrane, one on each side. Gill 
openings extending down to below the pre- 
opercular border, the distance between their 
lower ends equal to or less than an eye diame- 
ter. Ten or more developed gill rakers on the 
lower limb of the outer arch. Scales ctenoid, 
those on the body not especially enlarged 
posteriorly. Scales on top of head extending 
forward nearly to eyes, but no enlarged scales 
just behind eyes. First dorsal fin with 6 spines. 
First spine in both dorsal fins with a flexible 
tip. Caudal rounded, not longer than the head 
length. Pectoral without silky rays above. The 
united pelvics without a basal frenum, the 
membranes uniting the inner rays easily torn 
(as it apparently was in the holotype of 
Q. eugenius). 
Examination of numerous specimens of 2- 
eugenius^ 15 to 45 mm. in standard length, 
indicates no great sexual dimorphism. Over 
the available size range there is also no great 
differentiation in sensory papilla pattern. 
However, the cheek squamation does not ap- 
pear until well after all of the other scales 
have formed. 
67 
