68 
PACIFIC SCIENCE, VoL XIII, January 1959 
Among Hawaiian gobies Zonogobius (as 
represented by Z. farcimen) seems to be the 
closest relative. These two genera are set off 
from other Hawaiian gobies by the following 
features: no pelvic frenum; two longitudinal 
series of papillae on the flesh covering the 
premaxillary pedicels; and 10 or more de- 
veloped gill rakers. 
Of the three Hawaiian species of Quis- 
quilius, Q. eugenius is an abundant shallow- 
water flsh inhabiting areas of dead coral. It 
seems to be restricted to depths of less than 
15 feet. The other two species are in general 
deeper- water forms, though the type of Q. 
aureoviridis was collected in a tide pool with 
a maximum depth of 7 feet. 
Quisquilius aureoviridis sp. nov. 
Fig. 1, Table 1 
holotype: USNM 175013, 37.1 mm. in 
standard length taken in a rotenone station at 
Pupukea, Oahu, Territory of Hawaii by 
Gosline, et aL, Dec. 23, 1949. 
PARATYPES: USNM 175014, 1 specimen. 
36.2 mm., about 1 mi. north of Kailua, 
Hawaii, T. H., Gosline, June 19, 1953; 
University of Hawaii No. 1703, 8 specimens, 
21.0-27.2 mm., off Waikiki reef, Oahu, T. H., 
Gosline, et aL, Dec. 31, 1952. 
Depth of body contained about 4 times in 
the standard length. 
Head somewhat wider than deep, its great- 
est depth contained about 1.5 times in the 
head length. Mouth oblique, the lower jaw 
protruding. Front of mouth about on a level 
with the middle of pupil; rear of jaw about 
reaching to below front border of pupil. Eye 
bordered by a shallow groove posterodor- 
sally, its diameter contained about 4 times in 
head in a 2-inch specimen. Least distance be- 
tween eye and groove behind upper lip con- 
tained a little over 2 times in the eye diameter. 
Anterior nostril placed just above the groove 
behind upper lip, the posterior above the 
level of the top of pupil. Interorbital gently 
concave, not in a deep trench between the 
eyeballs, its width contained about 4 times in 
the eye diameter. Thirteen moderate, pecti- 
nate gill rakers on the lower limb of the outer 
arch in one specimen. Tongue rounded but 
usually with a central notch at tip. 
Body completely scaled, those scales above 
the pectoral base rather irregularly placed. A 
few scattered rows of papillae crossing certain 
of the body scales. Head scaled forward to the 
groove behind each eye. No scales on cheek 
or operculum. Thirty to 33 scales in a longi- 
tudinal series, 9 or 10 in a transverse series; 
about 13 predorsal scales. 
Some of the spines in the first dorsal fila- 
mentous. Second dorsal with a spine and 10 
or 11 soft rays. Anal with a spine and 8 or 9 
soft rays. Pectoral with about 19 rays. 
TABLE 1 
Certain Counts in Three Hawaiian Species of Quisquilius 
SPECIES 
LATERAL LINE 
SCALES* 
TRANSVERSE 
SCALE ROWSf 
TOTAL 2ND 
DORSAL RAYS 
TOTAL 
ANAL RAYS 
26 27 28 29 30 31 32 33 
7 8 9 10 11 
10 11 12 13 
9 10 11 
Q. eugenius 
1 1 3 
2 2 1 
4 1 
1 3 1 
Q. aureoviridis 
Holotype 
1 
1 
1 
1 
Paratypes 
111 1 
3 2 
2 3 
3 2 
Q. limhatosquamis 
Holotype 
1 
1 
1 
1 
Paratypes 
2 4 
3 3 
6 
5 1 
* Counted from the upper angle of the gill opening to the crease at the end of the hypural fan. 
t Counted from the beginning of the anal upward and forward to the dorsal base. 
