Alpheid Shrimp — BANNER 
143 
stead, it should read: "Dactylus simple ... as 
long as merus is broad.” 
Alpheus lanceloti Coutiere 
Fig. ^a-k 
For synonymy, see Banner: 1958: 167. 
LOCALITIES: 13 specimens, some ovigerous; 
Ahoolaka (”Sand Island”) Kaneohe Bay, 
Oahu, Hawaii; collected from silty sand flats 
at tide level by A. H. Banner. Maximum 
length about 20 mm. 
DISCUSSION: These specimens agree well 
with the original description by Coutiere on 
the general configuration and most details; 
there are, however, some differences. The 
rostral front in general agrees with Coutiere’s 
description and figures (1905^?: 900, pi. 63, 
fig. 39) but some variation occurs: the rostrum 
may be slightly longer, the rounded rostral 
ridge may not reach any further back than the 
middle of the orbital hoods, the frontal mar- 
gin at either side of the rostrum may be defi- 
nitely concave instead of almost straight, and 
finally the line or ridge at the inner margin of 
the orbital hoods is usually lacking. (”Le 
rostre est . . . separe des voutes orbitaires par 
un sillion. . . .”) Slight and variable differ- 
ences are similarly found in the proportions 
of the antennular peduncle and thoracic legs. 
In the large chela, as in the specimen already 
reported from Onotoa, the merus bears mov- 
able spines instead of teeth as shown by 
Coutiere; moreover, the ischium of the same 
appendage bears three or four movable spines. 
The greatest difference is in the small chela 
of the male, which Coutiere shows to be 
slender, with the fingers equalling the palm 
in length, while in these specimens the fingers 
are 1.2 times as long as the palm, slightly 
expanded in the middle, hooked on the end, 
and bear a fringe of setae, similar to a 
”balaeniceps-form,” except that it does not 
cross over the superior face of the chela. The 
small chela of the females and immature males 
is similar to that drawn by Coutiere. Coutike 
may have overlooked the sexual dimorphism. 
The small chela of the male from Onotoa was 
similar to the form described by Coutiere. 
It should be noted that the large chela is 
laterally compressed, approaching the condi- 
tion found in the Rapax group, but twisted, 
so that in these figures, and presumably those 
of Coutiere, the compression is not shown. 
In life the specimens had a basic light 
”sandy color,” similar in hue to their environ- 
ment; the chromatophores on their abdomens 
were in irregular and imperfectly defined 
bands. Their eggs were bright green. 
These specimens were found burrowing 
into the coherent fine sand of a broad sand 
flat. While their burrows were not traced, 
from the depths of the excavations necessary 
to capture the shrimp it is estimated that they 
penetrate into the sand about 12-15 inches. 
In the same environment, but burrowing more 
deeply, are found A. rapax (Fabricius) and a 
callianassid shrimp. 
The figures are included to complete the 
series of figures of Hawaiian species. 
Brevirostris Group 
Alpheus rapax Fabricius 
Figs. 9a-d, 103 
For synonymy, see Banner, Pacific Sci. 10(3): 
358, 1956. 
LOCALITIES: About 40 specimens, Kaneohe 
Bay, Oahu, in shallow burrows on the in- 
shore mud flats, intertidal zone; two speci- 
mens from deep sand burrows on Ahoolaka 
("Sand Island”) low in the intertidal zone, 
Kaneohe Bay, Oahu, Hawaii; collected by 
A. H. Banner, 1955, 1956. 
DISCUSSION: These specimens seem to be 
of two different size ranges according to their 
habitats; those from the shallow burrows 
(possibly up to 6 inches deep in the mud flats, 
with A. platyunguiadatus (Banner) occupying 
deeper burrows reaching into buried coral 
heads) were mature individuals with the 
carapace lengths up to 7.6 mm., while the two 
from the deep burrows in the fine sand of 
Ahoolaka were found from 12 to 18 inches 
down (with A. lanceoloti CoutiHe in shallow 
