Alpheid Shrimp — BANNER 
147 
of the ultimate article (Fig. I0a) \ in A. rapax 
this article bears a smaller group, not a tuft, 
of hairs, the longest of which reaches to the 
end of the ultimate article. In a few specimens 
of A. platyimgukulatus the tuft is not fully 
developed, but all lack the long setae of 
A, rapax. 
Alpheus platyungukulatus was separated from 
related species, especially from A. sauvensis de 
Man, on the basis of a series of minor differ- 
ences in the proportions of the appendages.^ 
This series of specimens was closely examined 
in those points of distinction, the results of 
which are given below: 
TYPE 
SPECIMEN 
RANGE OF 
THIS SERIES 
Ratio of lengths of first 
two antennular articles . . . 
1:1.3 
1:1.4-1:2.0 
Large cheliped 
Merus, length -4- breadth . . 
2.5 
2. 2-2.8 
Number of meral spines. . 
2 
2-5 
Chela, length breadth . . . 
2.3 
2. 1-2.6 
Chela, finger length 
palmar length 
0.5 
0.4-0.7 
Small Cheliped 
Merus, length -f- breadth . . 
2.6 
2.0-2. 8 
Spines 
4 
2-5 
Chela 
Palm, length breadth . . . 
1.5 female 
1.5 sole male 
Finger length 
palmar length 
1.6 female 
1.2 male 
1. 3-1.6 
females 
1.25 male 
Second legs, ratio of first 
two carpal articles 
1.2 female 
1:2.3 
1.25-1.50 
females 
1:1.85-2.3 
Third legs 
Merus, length breadth . . 
4.0 
4.3-5.0 
This range of variation destroys the value 
of most of the criteria set forth for the separa- 
tion of A. sauvensis and A. platyungukulatus; it 
also casts some doubts on the value of 
couplets used in de Man’s key to this group. 
However, de Man (1908:110) specified that 
^ In the tabulation on p. 132 of the original descrip- 
tion the reference to the stylocerite should be to the 
scaphocerite, and in the comparison to A. djeddensh in 
the paragraph preceding the tabulation the text should 
read "small chela’’ instead of "third chela.’’ 
the dactyls of the third to fifth legs in his 
species were simple and rounded, and in all 
these specimens they are definitely flattened. 
Moreover, de Man did not describe the tuft of 
dense hairs on the penultimate joint of the 
third maxillipeds, and he did describe the 
terminal setae of the last article of this ap- 
pendage as being over twice as long as the 
article, while in this species they are not even 
as long as the article. I believe, therefore, that 
this species is valid. 
Edwardsii Group 
Alpheus bastardi Coutiere 
Alpheus bouvkri var. hastardi Coutiere, Soc. 
Ent. de Erance, BuL, 1898(5): 133, fig. 
la, 1898. 
Alpheus hastardi Coutiere, Fauna and Geog., 
Maidive and Laccadive Archipelagoes 2(4) : 
907, pL 85, fig. 45, 1905. 
LOCALITIES: 1 specimen, Station Y-111, 
Tomil Harbor, Yap Island, Caroline Archi- 
pelago, from holes in intertidal zone, rocky 
beach; collected by R. W. Hiatt, 1946. 
DISCUSSION: This single specimen, a female, 
agrees perfectly with the short description and 
figures of Courtiere. Unfortunately there is no 
male with which to confirm the identification. 
Alpheus crassimanus Heller 
Fig. 11^-^ 
For synonymy, see Banner, Pacific Sci. 11(2): 
204, 1957. 
LOCALITIES: 3 specimens from Station Y- 
111, collection data as above. One specimen. 
Station 2015, Otikaheru Island, Tuamotu Ar- 
chipelago, on coral stones, under surface of 
muddy sand, near shore of brakish enclosed 
lagoon; collected by J. E. P. Morrison, July 
29, 1952. 
DISCUSSION: Most of the previous studies 
on this species have indicated that the small 
chela of the male not only has the dactylus 
subspatulate and fringed with long setae, but 
also that the palm bears sculpturing (see de 
Man, 1911, p. 417; Bate, 1888: pi. 99, Eg. 2; 
