180 
PACIFIC SCIENCE, Voi. XIII, April, 1959 
the stiff hairs 1.5-2 mm. long, white; stigmas 
briefly short exserted (for 1-1.3 mm.), low 
rounded, with a central crease; ovary 1.5 mm. 
long, ellipsoid; berry 6-7 mm. in diameter, 
globose-turbinate, apophysate, greenish; 
seeds 1.4-1. 5 mm. long, 0.7-0. 9 mm. wide, 
0.2-0. 3 mm. thick, elliptic, flat, with strong 
wavy parallel transverse ridges, the eminences 
rounded or muriculate, the raphe forming a 
prominent, thickened, straw-colored margin, 
the flat faces dull brown, but the color in 
places masked by the pale epidermis. 
HOLOTYPUS: Niihau, IF. T. Brigham (dis- 
tributed by Mann & Brigham), (BISH). 
SPECIMENS EXAMINED: Kauai ou Nihau 
( = Niihau), 1851-55, /. Remy 300 bis (GH; 
and photo in BISH). 
H. Mann, Jr., commented (1867: 180) on 
this Remy collection, “more probably Niihau 
— where it was also found by Mr. W. T. 
Brigham.” Rock (1919^/ 357) observed that, 
“The plants from Niihau collected by Remy 
are much more robust, the stems being nearly 
5 cm. in diameter.” The three sheets of the 
type collection, by Brigham, in the Bishop 
Museum, contain seven branches, all of them 
sections split longitudinally from the heavy 
and densely flowered stems. The longitudinal 
stem sectors were flattened in pressing, so 
that one cannot be sure of their original 
diam.eter. A revised estimate is 1-2 cm. at the 
base of the lowest leaves. 
The occurrence of a Delissea on Niihau is 
very noteworthy, for the lobelias in general 
and all of the species of Delissea in particular, 
occur in moist forests, usually in the rain- 
forest. That a Delissea was twice collected be- 
tween 1850 and 1865 by different botanists is 
an indication of the occurrence then on the 
uplands of a moist forest. Brigham, then the 
director of the Bishop Museum, is quoted by 
Forbes (1913: 25) as saying, “this is the only 
lobelia that he saw on the island, and that it 
was more plentiful over the area where it 
occurred than perhaps any other lobeliaceous 
plant occurring in an equal area on the [Ha- 
waiian] group.” The species is certainly ex- 
tinct now, having vanished when domestic 
grazing animals destroyed the native forest. 
Delissea undulata, collected by Gaudichaud 
in 1819 was described by him (1829: 457 and 
pL 78). His description in Latin consists of 
12 words, only 9 of which are descriptive. 
His own type specimen, now in Paris, gives 
the locality lies Sandwich. No more precise 
localization has been found. A photograph 
of his type specimen has been published by 
Rock ( 1919 : 201) and this shows a specimen 
in fruit and flower, and with slender, linear- 
lanceolate blades, coarsely and irregularly 
sinuate dentate. Though with broader leaves, 
a collection by Hillebrand, August 1870, dry 
pali of Olualu (Olowalu) gulch, West Maui, 
was photographed by Rock when in Berlin 
and this excellent photograph is in the Bishop 
Museum. It is similar to the generalized 
drawing published by Gaudichaud (1826-30, 
Atlas: pL 78). The plant has broader, less 
deeply lobed blades, but it appears to be of 
the same species, D. undulata Gaud., and it 
was so determined by Rock. 
There is a D. undulata Gaud. var. serrulata 
Wawra, described from fruiting material, 
Wawra 1,943, from Waihee, Maui. Rock 
( 1919 : 353) reduced this to the synonymy of 
the species and cited Wawra's specimen 
which he had studied in Vienna. This may 
well belong in the synonymy of D. undulata, 
but in lack of flowers it cannot be placed with 
certainty. It appears, then, that the D. undu- 
lata Gaud, was probably collected by Gaudi- 
chaud in the adjacent mountains when the 
“Uranie” was anchored at Lahaina, Maui. 
D. undulata Gaud, has the blades linear- 
lanceolate, coarsely and irregularly sinuate- 
dentate, and apparently much larger, while 
D. niihauensis has the blades broadly ovate, 
crenate, and 5. 5-7. 5 cm. long. 
D. fallax Hbd., apparently the closest rela- 
tive, has the blades 7-23 cm. long, 2. 3-5. 8 
cm. wide, narrowly elliptic, apiculate, ser- 
rate; petioles 3-13 cm. long; calyx lobes 
subulate, glabrous; corolla 15-17 mm. long, 
with a single, inconspicuous dorsal knob; 
