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PACIFIC SCIENCE, Vol. XIII, April, 1959 
parallel veins; ray flowers marginal, 6-9, 
bright yellow, the ray 7-9 mm. long, broadly 
elliptic, with numerous longitudinal parallel 
nerves, at apex emarginate and more or less 
3-toothed; corolla tube 1 mm. long; pappus 
awns unequal, 3-5, and 1-2 mm. long, linear 
or oblance-linear, thick, rigid, ascending sca- 
berulous, the larger ones channelled on the 
inner side; style 2 mm. long, filiform, the base 
not distended; stigmas 2, ligulate, 1.5-2 mm. 
long; the young ray achene 1.8-2 mm. long, 
1.7-1. 9 mm. broad, trigonous, and the broad 
distal face convex, broad cuneiform, the apex 
densely short white hispidulous; when ma- 
ture the ray achenes 2. 7-3.1 mm. long, 1.9- 
2.7 mm. wide, almost as thick, trigonous, 
broadly cuneiform, the sides brown, mottled 
with black and strongly tuberculate, the 
broad apex flatish or sloping, yellowish his- 
pidulous, awns 1.5-2 mm. long, subulate, 
ascending hispidulous, several of them per- 
sistent; disc flowers about 80; disc corollas 
4 mm. long, yellow, the tube 1.5 mm. long, 
tubular, the limb narrow funnel-form, 2.5 
mm. long, 5-lobed, the lobes 0.8 mm. long, 
deltoid, spreading; filaments 1 mm. long, 
filiform, pale; staminal tube 1.8 mm. long, 
0.5 mm. in diameter at apex, narrowing be- 
low, the linear anther sacs black, the apical 
connective tips pale, deltoid-ovate; style 3 
mm. long, pale, springing from a dark firm 
style base 0.4-0. 5 mm. long, cylindric, slightly 
tapering below, and with an expanded apical 
marginal rim; the 2 stigmas 1 mm. long, 
ligulate, acute; pappus of a single ligulate, 
subulate awn 2. 1-2.3 mm. long, attached 
apically near the base of the corolla tube; the 
young disk achenes 1.8-2 mm. long, 0.7- 
0.9 mm. wide at apex, narrowly cuneiform 
and tapering downwards, compressed, the 
two sides convex, glabrous, the sloping apex 
densely hispidulous; disk achenes when ma- 
ture 2.5 mm. long, 1.7-2 mm. wide, broadly 
cuneiform, yellowish, mottled with pale 
brown, partly smooth, partly low tuberculate, 
less thick than wide, but clearly tetragonal, 
the base obtuse or truncate, the apex broad. 
gently sloping, closely yellowish hispidulous, 
the margin with low tubercles, the usually 
single awn apical, 1-2.3 mm. long, subulate, 
ascending hispidulous, sharply bent. 
HOLOTYPUS: Niihau, Kawaihoa Point, 300 
ft. alt., in dry tuff, head of steep gully, erect, 
much branched shrubs, rays yellow, disk dark 
yellow, March 31, 1949, H. Sl John 25,611 
(BISH). 
This plant has been carefully compared 
with the known species and varieties of 
Lipochaeta. It is outstanding in the genus by 
having the receptacle at first lanceoloid, then 
ultimately conic, instead of flat or gently 
convex. It is most similar to L. lobata var. 
denticidata of the island of Oahu, but this va- 
riety has the petioles 2-5 (-9) mm. long; inner 
and outer phyllaries oval, subequal; chaff 
obtuse; ligulate flowers 8-18; and the achenes 
2. 9-3. 6 mm. long, verrucose, sharply angu- 
late, narrowly obdeltoid, compressed. L. ka- 
waihoaensis differs from it by having the 
petioles 4-18 mm. long; the inner phyllaries 
lanceolate, longer and narrower than the 
outer ones; chaff acute; ligulate florets 6-9; 
and the achenes 2-3.1 mm. long, 1.7-2. 7 
mm. wide, the sides tuberculate, the angles 
not winged or margined. There is a lesser 
similarity in appearance to L. tenuis var. 
Sellingii of Puu Hapapa, Oahu, which variety 
is different in having the habit decumbent or 
suberect; blades oblong-lanceolate, at base 
broadly cuneate, the margin sharply serrate; 
peduncles 2-5 cm. long; disk flowers with 
corolla lobes equaling the tube, the pappus 
of 4 bristles as long as the corolla tube, and 
the achenes obovate, triangular with the an- 
gles more or less winged. On the other hand, 
L. kawaihoaensis can be known by having the 
habit erect; blades ovate to deltoid-ovate, at 
base rounded, the margin coarsely crenate or 
even serrate; peduncle 3-17 cm. long; disk 
flowers with corolla lobes Vi as long as the 
tube, pappus of a single ligulate-subulate 
awn slightly more than as long as the 
corolla tube; achenes broadly cuneiform, 
tuberculate, not winged. 
