188 
PACIFIC SCIENCE, VoL XIII, April, 1959 
brous, and white subappressed hispidulous, 
broadly ovate, the base rounded or subcor- 
date, the apex abruptly subacute, the margin 
doubly dentate-serrate, below a little paler, 
triplinerved from just above the base; heads 
in a loose, mostly 3-headed terminal cyme, 
or with 3-headed cymes on weak, late, lateral 
branchlets, peduncles 1.5-6. 5 cm. long, naked, 
closely subappressed white hispidulous; heads 
heterogamous, radiate; involucre in anthesis 
4-5 mm. tall, 8-10 mm. in diameter, but in 
fruit more widely spreading, knee-pan-shaped ; 
phyllaries in 2 (or 3) rows, greenish, the outer 
ones 3-3.5 mm. wide, 5 (-7) in number, 
broadly ovate, subacute, appressed hirsutu- 
lous on the outer surface and the margin, firm 
coriaceous except at the firm fleshy base, 7- 
nerved; inner phyllaries slightly longer, 2.5 
mm. wide, obovate, acute; receptacle in 
anthesis low conical, 1 mm. tall, 2 mm. broad; 
chaff 3.5-5 mm. long, narrowly wedge- 
shaped, the apex rounded to an acute tip, 
folded along the midrib, the sides 0.8-1 mm. 
wide, many nerved, scarious, the exposed tips 
hispidulous; ray flowers marginal, 8-14, when 
fresh yellow, the ray 8-9 mm. long, 3.8-4 mm. 
wide, broadly oval, shallowly 3-toothed at 
apex, on the distal side with 2 strong salient 
ribs from which the sides are infolded, weaker 
longitidunal ribs 8; ray corolla tube 1.4 mm. 
long; pappus awns mostly 3, unequal, 0.6- 
0.9 mm. long, subulate, ascending scaberu- 
lous; styles 2.5 mm. long; stigmas 0.7 mm. 
long, filiform; the young ray achenes 1.2-1. 4 
mm. long, trigonous cuneiform, the margins 
and apex hispidulous ciliate; when mature the 
ray achenes 2. 5-2. 8 mm. long, 1.2-1. 8 mm. 
wide and nearly as thick, sharply trigonous, 
broadly cuneiform, dark brown, mottled with 
gray, towards the apex rough hispidulous and 
less so towards the base, the angles lacerate 
winged, awns 0.6-1. 2 mm. long, 3 (or 4), 
unequal, subulate; disk flowers about 100; 
disk corollas when fresh greenish yellow, 4 
mm. long, the tube 1 mm. long, cylindric, the 
limb very narrowly funnel-form, 0.9 mm. in 
diameter, 4-lobed, the lobes 0. 5-0.6 mm. 
long, deltoid, ascending; filaments 1 mm. 
long, ligulate, pale; staminal tube 1.4 mm. 
long, 0.5 mm. in diameter, tapering down- 
wards, the anther sacs blackish, the apical 
connective tips pale, deltoid-ovate; style 3.5 
mm. long, filiform, pale, springing from a 
dark, cylindric style base 0.3 mm. long, with 
the upper rim expanded and lobed; the 2 
stigmas 0.7-0. 8 mm. long, ligulate with a 
narrowly rhombic tip; pappus of two subulate 
awns 0.7-1. 3 mm. long, upward barbellate, 
attached above the lateral angles; the young 
disk achenes 1.3-1. 9 mni. long, 0.6-0. 8 mm. 
wide, thin biconvex, wedge-shaped, hispidu- 
lous on margins and apex; when mature the 
disk achenes 2. 8-3. 2 mm. long, 1.1-1. 7 mm. 
wide, strongly compressed and biconvex, 
elliptic-obovate or narrowly so, the base ob- 
tuse, the apex truncate or excavate, the surface 
mottled brown and gray, the sides smooth 
towards the base, but upwards tuberculate 
hispidulous; each of the margins bearing a 
thin wing that is entire, lacerate, or hispidu- 
lous-ciliate lacerate; awns two 0.7-1. 7 mm. 
long, equal or unequal, subulate, upwardly 
barbellate, attached apically. 
HOLOTYPUS: Niihau, Kii, among rocks on 
basalt knoll, 100 ft. alt., April 2, 1949, H. Sl 
J ohn 23,664 (BISH). 
SPECIMENS EXAMINED: Nfihau, Kii, be- 
tween basalt boulders on rocky knoll, 75 ft. 
alt., April 2, 1949, H. Sl John 23,67 L 
L. niihauensis is a member of the section 
Lipochaeta {Microchaete ) . Its closest relative is 
L. subcordata Gray populifoUa Sherff of the 
island of Lanai, which variety is distinguished 
by having the stems 3-6 dm. tall; petioles 
1-3 cm. long, slender; blades from narrowly 
ovate-cordate to lanceolate-cordate or nar- 
rowly deltoid, the principal ones broadly 
ovate, subcordate, the margins doubly sharply 
attenuate serrate; outer phyllaries about 4 
mm. long; ray flowers 5-6, and 5-7 mm. long, 
their achenes 2-2.5 mm. long, 1.8-2 mm. 
wide, black, oblong-cuneate, trigonous, wing- 
less, the apex truncate, the back with many 
high tubercles; disk corollas 3 mm. long. 
