Flora of Papua New Guinea 
Lae National Herbarium 
9225 Polygalaceae 
/ Xanthophyllum papuanum Whitm. 
Southern Highlands province, Kutubu, limestone ridge S of Ubogo 
village, mature hill forest, 425-500 m elev., near 6° 33’ S, 143° 12’ E. 
with sequence starting 921 1 ; canopy tree on karst, outer trunkbark dark 
brown, thick, straw-colored, lenticels verrucose, inner bark furrowed, 
wood light brown, dense; branchlets slender, terete, shining green, 
glabrous; leaves alternate, distichous, spreading, coriaceous, glabrous, 
exstipulate, adaxially dark shiny green, abaxially slightly lighter green; 
blade lanceolate, typical 1 1 x 4.5 cm, apex acuminate, base cuneate; 
venation 3-5 lateral pairs, arcuate, not closing, reticulum irregular, 
obscure; petioles 0.5-1 .0 cm, flat above, glabrous, transversely lined and 
with a glandlike prominence in the axil; flowers not seen; infructescence 
terminal, rachis stout, woody, puberulent, marked by small abscission 
scars; fruits globse, ca 10 cm dia in vivo, heavy, exocarp dull whitish- 
green, smooth, glabrous, pericarp hard, splitting irregularly when mature, 
seeds numerous, flat on the commissural faces, covered by a yellow- 
orange arillode (seeds are reported by villagers to be eaten by hornbills 
and cassowaries). 
NCI screening code: U44Z-8498-W, -8499-X, -8500- Y, -8501 -Z, -8502-A, 
corresponding to TWD, RT+RTWD, TBK, LF, TW+BR; 6 herbarium 
duplicate sets distributed through NCI. 
carpological collection made. 
coll. W. Takeuchi 9225 
date: Sept. 13, 1993 
det WT 
Flora of Papua New Guinea 
Lae National Herbarium 
9235 Annonaceae 
Friesodielsia or Mitrella 
Southern Highlands Province; Kutubu district, swamp forest adjacent to 
Waro and Ubogo villages, NE towards the low ridge, elev. 425 m in 
Metroxylon-dominant swamp, canopy somewhat open, lat. 6 deg. 32’ S x 
long. 143 deg. 12’ E. 
w. 9228; woody climber into the canopy; twigs black, slender, terete, 
glabrous, rugulose (lens); leaves alternate, distichous, coriaceous, 
glabrous, exstipulate, adaxial surface dark dull green x abaxially light 
green; lamina linear-elliptic, typical blade 9 x 2.25 cm to 11 x 2.75 cm, 
apex acuminate, base rounded; venation pinnate and reticulate but only 
the midrib manifest; petiole 1 cm, channelled, glabrous; infructescence a 
terminal etaerio, glabrous, ebracteate; peduncle green, to 4 cm long; 
receptacle yellow-green, not expanded, subtended by the persistent calyx 
of 3 divaricate and triangular sepals; berry stipitate, the stalk 4-7 mm, 
both the berry and stalk dull orange; individual fruits typically misshapen, 
irregularly ovoid to oblongoid, to 20 x 12 mm; mesocarp orange; seeds 1- 
4, yellow and white ruminated; flowers extra-axillary, solitary; sepals 3, 
minute (1-2 mm), nearly free, puberulous; corolla biseriate, 3+3; the outer 
whorl contorted, valvate, 30 mm long, glabrescent outside, puberulent 
and matted inside, not clawed; inner petals 7-8 mm long, hairy at the tips 
(both inside and out_more so on the adaxial surface); stamens numerous, 
inserted on an elevated torus, anthers ?4, obscured by the expanded 
connective; gynoecium apocarpous, about 5-carpellate, the carpels 
oblong, glabrous, crowned by the clavate stigma. 
dissected flower, berry bottled, at A. 
coll. W. Takeuchi 9235 
date: Sept. 14, 1993 
det.: WT 
