Flora of Papua New Guinea 
Lae National Herbarium 
10119 Rhamnaceae 
Zizyphus angustifolius (Miq.) Hatusima 
East Sepik Province, lowland foothill forest on the ridge and slopes 
above Garuka village, near 4° 11’ S, 142° 44’ E. 
with 10118 near top of buttress ridge; elev. c. 300 m; 15 m tree in 
shallow draw below the crest; twigs slender, terete, orange-brown 
pubescent, apically somewhat fractiflex, branching pattern plane; 
leaves alternate, distichous, apparently minutely/caducously 
stipulate, lamina linear-elliptic or linear-lanceolate, papery, to 
approx. 23.5 cm x 7.5 cm, apex acuminate, margin distantly 
serrulate, base obtuse and often unequal, blade concolorously dull 
dark green on both surfaces, adaxially somewhat bullate; petiole c. 
1.5 cm, flattened above, puberulous; leaf venation triplinerved, 
costae connivent at both poles, impressed above, prominent 
beneath, plexus lax, irregular, the major intercostal nerves forming 
crossbars perpendicular to the longitudinal axis; fruit not seen; 
inflorescence axillary, abbreviated, 2 cm length, cymose, peduncle 
short, all axes puberulent, bracteolate; flowers congested, radial, 
dichlamydeous, the calyx green, 6-merous, gamosepalous with 
exterior surfaces and pedicels appressed hairy, sepals divaricate at 
anthesis, corniculate and with a median longitudinal ridge; stamens 
6, alternate with the sepals, obliquely spreading, enveloped by the 
petals; disk incrassate, fleshy, 6-lobate, each lobe with 2 channels; 
ovary inferior, stigmas 3. 
Waskuk name-mangana,’ (accent on the first syllable). The wood is 
used ip various constructions. 
coli: W. Takeuchi 10119 
det. WT 
date: August 22, 1 994 
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