Flora of Papua New Guinea 
Lae National Herbarium 
10356 Rutaceae 
Tetractomia cf. tetrandrum (Roxb.) Merrill 
East Sepik Province; April River at Pinape; pristine lowland hill forest 
on mafic substrate, ascending Crestline at elev. 350’ altimeter (c. 105 
m); colls, centered on 4° 33’S latitude x 142° 34’E longitude. 
w. collection sequence 10354-10364; small tree on steep slopes off 
the Crestline, single sighting; outer bark smooth, pale brown, wood light 
yellow, sap lacking; stipules 0; leaves opposite, glabrous, glandular- 
punctulate, fleshy, adaxial surface glossy medium green x abaxially 
glossy light green; lamina orbicular-obovate, typical blade 12 x 7.5 cm, 
apex rounded, margin entire, base cuneate; lateral venation in approx. 
8-10 strong pairs with weaker intervening nerves between, reticulum 
obscure; petiole swollen at both ends, geniculate at the blade, c. 3 cm 
long; infructescence axillary, highly ramifying, open, the axes 
bracteate, puberulent with both simple and stellate hairs; fruits 
immature, 4-lobed and light green when fresh, splitting into 4 
schizocarp-like members after drying; epicarp glabrous, glanduliferous; 
endocarp pergamentaceous and separating; ovule apparently 1 in 
each cell (near the base), minute; perianth persisting on the developing 
fruit, 4-merous; staminal residue 4, eciliate, aglandular, egeniculate; 
one style remnant seen with a small bilobate stigma. 
tokples name (Bugabugi) = ‘oh say;’ wood is used for houses and 
flooring. 
NCI screening code: OFCZ-0808-N, -0809-0, -0810-P, -081 1-Q, - 
0812-R, -081 3-S; corresponding to LF+TW+WD, TBK+sapwood, 
FL+RTWD, RT, TWD, BR; 6 herbarium duplicates distributed through 
NCI. 
coll: W. Takeuchi 10356 
date: July 21, 1995 
det: P. F. Stevens, Feb. 15, 1996 
Flora of Papua New Guinea 
Lae National Herbarium 
10357 Verbenaceae 
Teijsmanniodendron ahernianum (Merr.) Bakh. 
East Sepik Province; April River at Pinape; pristine lowland hill forest 
on mafic substrate, ascending Crestline at elev. 400’ altimeter (c. 122 
m); colls, centered on 4° 33’S latitude x 142° 34’E longitude. 
c. 15 m pole tree, common species; bole straight and unbuttressed; 
twigs apically puberulent; leaves opposite, digitately trifoliolate, firm, 
glabrous, medium shiny green both surfaces; inflorescence terminal, 
paniculate, ascending, peduncled, bracteate, all parts with a coarse 
puberulence; calyx 5-dentate, hairy, slightly accrescent; fruits 
indehiscent, epicarp glossy black and glabrous; seed 1 . 
Bugabugi name = 'me mape.’ A strong timber used in construction. 
NCI screening code: OFCZ-0814-T, -0815-U, -0816-V, -0817-W, - 
081 8-X, -081 9-Y; corresponding to LF+TW+BR, RT+RTBK, BR, TBK, 
TWD, RTWD; 8 herbarium duplicates distributed through NCI. 
coll. W. Takeuchi 10357 
date: July 21, 1995 
det.: WT 
