The Nature Conservancy Botanical Survey 
Flora of Josephstaal, Papua New Guinea 
Lae National Herbarium 
13412 Thelypteridaceae 
Pronephrium micropinnatum Holttum 
keys to this species on Holttum 
Madang Province, Josephstaal FMA area, streambed flowing 
to SW of expedition Camp 1 (‘Kumamdeber’), centered on 
GPS 281407 E, 9504560 N, elev. 530’ (160 m), mature 
growth foothill forest with occasional landslip communities. 
coll, sequence 13400-13436; saxicolous herbs on streambed; 
stipes black; leaves herbaceous, aglandular, upper side very 
dark green, medium green beneath, indument of setiform 
hairs; sori exindusiate, not confluent; sporangia setiferous 
only. 
coll.: W. Takeuchi, J. Wiakabu, M. Gorrez, A. Towati 13412 
date: July 28, 1999 
det: WT; Jan. 4, 2000 
