64 
SALIX 
Messrs Marshall and Shoolbred ( fourn . Bot. xlvii, 222 (1909)) record a Sutherlandshire plant which the Rev. E. F. Linton 
suggested had the above parentage. We do not doubt that such complicated hybrids, and even hybrids still more complicated, 
occur in nature ; but it appears to us that the results of very precise and very numerous experiments on hybrids by recent 
Mendelian workers have established beyond doubt that it is not possible to discover precisely the parents of such putative 
hybrids by morphological methods. The same parentage has also been ascribed to a plant collected in Germany.] 
5 . caprea x viminalis , although local, is rather widespread in England, rather rare in eastern Scotland, and 
“not unfrequent in Ireland” (Syme, under x S. smithiana, op. cit. p. 227); from Cornwall, Sussex, and Essex 
northwards to Perthshire and Ross-shire ; co. Cork, co. Kildare. 
Southern Scandinavia, Denmark, Germany, Holland, Belgium, France, central Europe, Russia ; Asia eastwards 
to Japan. 
S. cinerea x viminalis Wimmer in Flora xxviii, 437 (1845) emend.; in Flora xxxi, 318 (1848); in 
Denkschr. Schles. Gesellsch. 161 (1853) including 5 . dasyclados x viminalis p. 160, 162; Sal. Eur. 181 (1866) in- 
cluding xS. stipularis p. 184 et x S. holosericea p. 189; A. et G. Camus Classif. Saul. 314 (1904) including 
x S. stipularis p. 318, excluding syn. Forbes et syn. Koch and their equivalents; v. Seemen in Ascherson 
und Graebner Syn. iv, 266 (1909) including 5 . dasyclados subsp. stipularis p. 180, excluding syn. 5 . ferruginea 
Forbes et 5 . smithiana Forbes et syn. Leefe ; Rouy FI. France xii, 237 (1910). 
leones: — Forbes Sal. Woburn, t. 129, as S. geminata\ t. 135, as S', micheliana. 
Exsiccata : — E. F. et W. R. Linton, 10; Schultz x, 921 ; Wimmer et Krause ( H . SI), 24. 
Small tree or shrub. Young branches long, rather stout and coarse, persistently pubescent. 
Stipules caducous or persistent, very variable in size and shape, often large and sometimes stalked 
and serrate on the summer shoots and coppiced shoots. Petioles about ro — 1*5 cm. long, pubescent. 
Laminae lanceolate, broadly lanceolate, or narrowly obovate, usually narrower than in S. caprea x 
viminalis , margin often somewhat incurved and crenulate, acute to acuminate or gradually tapering 
to the apex, covered with persistent hairs on both surfaces, very hairy underneath, from about 
8 — 16 cm. long or rather longer, and 1 ’5 to 3*0 broad. Catkins smaller than those of .S’, caprea 
x viminalis, larger than those of S. aurita x viminalis ; April. Bracts as in these hybrids. Style 
and stigmas variable, but usually long. Capsules pubescent, stalked ; May. 
(B) x S. holosericea Wimmer Sal. E?ir. 189 (1866); A. et G. Camus Classif. Saul. 314 (1904); v. Seemen in 
Ascherson und Graebner Syn. iv, 267 (1909); 5 . holosericea Willdenow Berl. Baumz. 458 (1796) ; Hooker Brit. FI. 
ed. 4, 364 (1838); x S. lancifolia Doll FI. Baden. 512 (1859); x S. smithiana var. velutina 1 Andersson in DC. 
Prodr. xvi, pt. ii, 268 (1868) excl. var. ferruginea ; White in fourn. Linn. Soc. xxvii, 418 (1890). 
leones: — Reichenbach Icon. t. 579, fig. 2026 [1226], as 5 . holosericea ; Hartig Forst. Cult. t. 112 (37b), as 
S. holosericea ; A. et G. Camus op. cit. t. 29, fig. M — R, as x S', holosericea. 
Exsiccata : — Fries, xi, 61, as S. holosericea. 
Stipules, when persistent, smaller than in x S. stipularis. Laminae shorter, up to about 
8 - o cm. long and 1*5 broad, with grey or rust-coloured hairs underneath. Catkins smaller than in 
x S’, stipularis. Style rather short but distinct. Stigmas entire or bifid. 
(C) x S. stipularis A. Kerner in Verhl. Z.-B. Gesellsch. Wien (217) (i860); Wimmer Sal. Eur. 184 (1866); 
A. et G. Camus Classif. Saul. 318 (1904); S. stipularis Smith Eng. Bot. no. 1214 (1803)!; FI. Brit. 1069 (1804); 
Eng. FI. iv, 230 (1828); Syme Eng. Bot. viii, 225 (1868); S. viminalis x dasyclados Wimmer in Denkschr. Schles. 
Gesellsch. 160 (1853); x S', smithiana var. stipularis White in fourn. Linn. Soc. xxvii, 415 (1890); S. dasyclados 
subsp. stipularis Ascherson und Graebner Syn. iv, 180 (1909); S. cinerea x viminalis race stipularis Rouy FI. 
France xii, 238 (1910). 
leones : — Smith Eng. Bot. t. 1214 (pistillate catkins immature, and leaves from summer shoots), as S', stipularis-, 
Forbes Sal. Woburn, t. 130, t. 132, as S. stipularis-, FI. Dan. t. 2268, as S. stipularis-, Reichenbach Icon. t. 598, 
fig. 1249, as S. stipularis ; A. et G. Camus op. cit., Atlas t. 29, fig. J — K, as x S. stipularis. 
Exsiccata: — Leefe, i, 15, as S. stipularis ; E. F. et W. R. Linton, 9, 84, as S. stipularis. 
Stipules often caducous on the normal leaves ; those of the summer shoots persistent, more 
or less stalked, large, long, more or less coarsely serrate on the outer margin, often with a 
large tooth at the base, acute, pubescent underneath. Laminae longer and relatively more narrow 
than in xS. holosericea , up to about 17 cm. long and 2‘5 — 3 - o broad, grey or white with hairs 
underneath. Catkins longer than those of x S. acuminata or x .S’. smithiana. Style variable in 
length. Stigmas linear, divided or not. Staminate plants appear to be rare. 
1 The name “ S. velutina Schrader” would appear to be illegitimate. It seems to be based merely on a citation in 
synonymy by Koch (Syn. 650 (1837)) as follows: — “ S. velutina Schrader secunda specimina ex horto Gottingensi in herbario 
Mertensio.” 
