ULMUS 
89 
bracts destitute of flowers, appearing before the leaves. Perianth campanulate, persistent, with 4 — 9, 
usually 4 — 5 divisions. Anthers reddish before dehiscence. Ovary usually unilocular, compressed. 
Stigmas 2. Fruit a winged achene, i.e., a samara. Wing broad, green, more or less notched 
at the apex. 
About 20 species ; north temperate zone, mountains of tropical Asia. 
The British species belong to the subgenus Dryoptelea (Spach in Ann. Sci. Nat. ser. 2, xv, 361 (1841); Engler in 
Pfianzenfam. iii, pt. i, 62 (1894)). 
British series of Ulmus 
Series i. Nitentes (see below). Petioles long or rather long. Laminae of normal leaves smooth 
above at maturity; of the suckers, adventitious shoots, and of the summer-shoots rough above. 
Fruit usually obovate. Seed placed between the middle of the fruit and the apical notch. 
Series ii. Campestres (p. 94). Petioles rather long. Laminae of all the leaves rough 
above. Fruit small, suborbicular. Seed placed as in Nitentes. 
Series iii. Glabrae (p. 95). Petioles short or very short. Laminae of all the leaves very 
rough above. Fruit large, elliptical to obovate. Seed placed in the centre of the fruit. 
Series i. Nitentes 
Nitentes nobis. 
For characters, see above. 
Species of Nitentes 
1. U. nitens (see below). Large tree. Lower branches wide-spreading. Laminae very unequal 
at the base, very smooth and shining above. Fruit obovate. 
U. glabra X nitens (p. 91). Large trees. Laminae larger than in U. nitens , usually smooth and shining 
above. Fruit larger than in U. nitens. 
2. fU. stricta (p. 92). Tree rather small, pyramidal. Branches short, more or less as- 
cending. Laminae not very unequal at the base, smaller than in U. nitens. Fruit as in U. nitens. 
3. U. sativa (p. 93). Tree rather small. Branches rather short, lower ones wide-spreading. 
Laminae not very unequal at the base, smaller than in U. nitens. Fruit smaller than in U. nitens, 
oblong-elliptical to obovate. 
I. ULMUS NITENS. Smooth-leaved Elm. Plates 90, 91, 92, 93; 94, 95, 96, 97 
Ulmus folio glabro Goodyer in Johnson’s Gerard Herb. ed. 2, 1481 (1636); Parkinson Theatr. Bot. 1403 
(1640); Ray Syn. ed. 3, 469 (1724). 
Ulmus nitens Moench Meth. Plant. 333 (1794) ; Moss in Gard. Chron. ser. 3, li, 199 et 217 (1912) ; U. glabra 
var. Hudson FI. Angl. 95 (1762); U. glabra Miller Gard. Diet. ed. 8, no. 4 (1768) non Hudson; Lindley Syn. 
226 (1829)!; U. campestris var. glabra Aiton Hort. Kew. i, 319 (1789); U. surculosa var. glabra Stokes Bot. Mat. 
Med. ii, 35 (1812); U. campestris var. laevis Spach in Ann. Sc. Nat. ser. 2, xv, 362 (1841); U. suberosa var. 
glabra Syme Eng. Bot. viii, 138 (1868); U. vulgaris 1 var. carpinifolia 2 Rouy FI. Fra7ice xii, 2 66 (1910); U. cam- 
pestris race glabra Ascherson und Graebner Syn. v, 553 (191 1) partim. 
leones: — FI. Dan. t. 632, as U. campestris ; Duhamel Traite des Arbres iii, t. 42, as U. campestris. 
Exsiccata : — Billot, 1763 (partim) as U. campestris ; Fries, viii, 57, as U. campestris var. glabra. 
Tree, attaining a height of about 30 — 35 m. Timber said to be valuable. Bark of old 
trees often striated by long oblique ridges and furrows. Branches large, lower ones wide-spreading, 
upper ones ascending, terminal ones frequently drooping. Young branches much more slender 
than in U. glabra or in U. campestris or in U. glabra x nitens, smooth during the first year, 
M. 11. 
1 U. vulgaris Pallas Reise iii, 135 (1776) is a name in a list and without any description. 
2 We have been unable to find the name U. carpinifolia in Ehrhart’s Beitrage. 
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