SA LSOLA 185 
either a succulent or membranous pericarp, enclosed in the winged and enlarged perianth. Seeds 
horizontal. Integument single. Embryo green, cochleate. Endosperm absent. 
About 40 species; Europe; temperate Asia; northern and southern Africa; chiefly in saline 
situations. 
British species of Salsola 
1. S. kali (see below). Usually much stouter than S. tragus. Spines of the leaves usually 
stronger. Wings of the fruiting perianth pronounced. Achene larger, about 2’ 5 mm. long and 3 ’5 
broad. 
2. # S. tragus (page 186). Stem slender. Leaves slender, about 2 — 5 cm. long, scarcely 
succulent. Wings usually absent, if present shorter than in S. kali. Achene smaller, about 2 mm. 
long and broad. 
I. SALSOLA KALI. Prickly Saltwort. Plates 192, 193, 194 
Kali Lyte New Herball 127 (1586); Tragos matthioli sen potius tragus improbus matthioli Gerard Herb. 959 
0597); Tragos sive tragurn matthioli Parkinson Theatr. Bot. 1034 (1640); Kali spinosum cochleatum Ray Syn. 
ed. 3, 159 (1724)- 
Salsola kali L. Sp. PI. 222 (1753)!; Miller Gard. Diet. ed. 8, no. 1 (1768)!; Smith Eng. Bot. no. 634 
0799); Ft. Brit. 280 (1800); Eng. Ft. ii, 18 (1824); Syme Eng. Bot. viii, 4 (1868); Rouy FI. France xii, 65 
(1910) excl. race gmelini. 
leones : — Svensk Bot. t. 471, as S', kali. 
Camb. Brit. FI. ii. Plate 192. {a) Flowering shoot of var. hirsuta. Norfolk (C. E. M.). (b) Flowering shoot 
of var. glabra. ( c ) Portion of stem of var. glabra. ( d ) Ripening ovary (enlarged). Sussex (T. H.). 
Annual. Root strong, penetrating the soil to a considerable depth. Stem erect, decumbent 
or prostrate, up to about 6 dm. high, though usually about half this height, with pale green 
or reddish stripes, usually much branched from the base. Branches spreading or ascending. 
Leaves sessile, succulent, subterete, subulate, often rather recurved, about 1 — 4 cm. long, attenuate 
at the apex into a little spine. Bracteoles 2, in the axils of the leaves, leaflike. Flowers 1 — 3 
in the axil of a leaf or leafy bract; opening in July. Perianth with 4 — 5, usually 5 segments; 
segments lanceolate, membranous during the flowering period, becoming more or less cartilaginous 
in fruit and markedly thickened about the middle, the thickening forming sometimes a mere 
ridge and at other times forming horizontally spreading wings of variable size. Stamens 3 — 5, 
usually 5. Anthers pale yellow. Style rather longer than the stigmas. Stigmas 2 — 3. Achene 
turbinate, about 2 ’5 mm. long and 3 - 5 broad, covered with the persistent perianth. 
The short leaved forms have been named var. brevifolia (Du Mortier FI. Belg. 23 (1827) nomen), and the longer- 
leaved forms var. longifolia (Du Mortier loc. cit. nomen = var. tenuifolia Reichenbach FI. Excurs. Germ. 583 (1832) non 
aliorum). Plants with stouter leaves have been named var. crassifolia (Reichenbach loc. cit. = var. latifolia Schur Pl. Transsilv. 
568 (1866)). Plants with rudimentary wings have been named var. marginata by Celakowsky (FI. Bohm. 155 (1867)). 
(a) S. kali var. hirsuta Hornemann Oec. Plant, ed. 3, i, 293 (1821); S. decumbens Lamarck Ft. France iii, 
241 (1778); S. kali var. hirta Tenore Syll. FI. Neap. 124 (1831); Rouy FI. France xii, 65 (1910); S. kali var. 
vulgaris Koch Syn. ed. 2, 693 (1844); S. kali var. typica Beck FI. Nied.-Ost. 340 (1890). 
leones: — Ft. Dan. t. 818 (left-hand plant), as S'. / kali-, Smith Eng. Bot. t. 634, as S'. kali\ Pallas III. t. 28, 
fig. 2, as S. kali\ FI. Lond. ed. 2, t. 158; Beck in Reichenbach Icon, xxiv, t. 292. 
Camb. Brit. FI. ii. Plate 193. {a) Upper portion of plant. ( b ) Portion of stem (enlarged). ( c ) Infructescence 
(enlarged). Sussex (T. H.). 
Exsiccata : — Billot, 841, as S. kali\ Dickson, xii, 14, as S. kali\ Hansen, 868; Magnier, 35, as S. kali\ 
Schultz, x, 904, as S. kali. The specimens by Billot and Schultz belong to the slender-leaved form. 
Stem prostrate or ascending, asperous. Leaves asperous. Wings of the mature perianth dilated 
or rarely rudimentary. 
This is the common British plant. 
Scandinavia, Denmark, Germany, Holland, P'rance, Italy, and doubtless elsewhere. 
(1 b ) S. kali var. glabra Detharding Consp. PI. Megalop. 25 (1828); Tenore Syll. FI. Neap. 124 (1831) excl. 
syn. L. ; S', spinosa Lamarck FI. France iii, 240 (1778) excl. syn. L. ; S. tragus DC. FI. France iii, 396 (1815) 
non Linn.; S. kali var. tragus Moquin in DC. Prodr. xiii, pt. ii, 187 (1849) excl. syn. L. ; Rouy FI. France xii, 
65 (1910) excl. syn. L. ; S. kali var. calvescens Grenier et Godron FI. France iii, 31 (1855). 
M. II. 
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