Alpheid Shrimp, X: Fiji, Tonga, Samoa — Banner and Banner 
161 
cerite with acute, narrow tip extending con- 
siderably beyond end of antennular peduncle; 
lateral margin curved, proximally provided near 
articulation with strong tooth at right angles 
to margin; squamous portion short and narrow 
reaching only slightly beyond end of second 
antennular article. Carpocerite equal in length 
to scaphocerite. 
Middle article of third maxilliped with strong 
rounded projection on inferior external margin. 
Merus of large cheliped less than 0.3 length 
of chela; superior margin projecting but not 
acute; inferior internal margin with several 
slight rounded protuberances and bearing short, 
soft setae and two short spines distally; article 
otherwise without setae except on superior distal 
margin. Chela heavy, compressed, 2.1 times as 
long as broad at maximum breadth, with fingers 
occupying only distal 0.2 of entire length. 
Palm with sharply delimited superior groove 
extending full length of chela; superior crest 
terminating in heavy rounded projection above 
dactylar articulation. Projection especially con- 
spicuous on inner face. Plaque crest flat, mar- 
gins sharp, terminating in two rounded teeth 
flanking adhesive plaque. Palmar groove shallow 
but well delimited distally. Inferior crest flat- 
tened, ending in small acute tooth. Inferior 
depression rounded. Dactylar articulation also 
flanked with strong tooth on superiomedial side. 
Dactylus heavy and rounded. Upper portion of 
superiomedial face papillose, papillae bearing 
short and mostly light setae; other portions of 
chela with at most scattered setae; chela not 
conspicuously hirsute. 
Small cheliped heavy. Superior margin of 
merus projecting distally into low rounded 
tooth; inferior internal margin armed with three 
small spines carried on heavy protuberances. 
Carpus with superior distal margin projecting 
as strong tooth over base of chela. Chela 3 
times as long as broad, with fingers occupying 
distal 0.45. Palm with superior groove and crest 
similar to those of large chela, except shallow 
and poorly developed; superior crest projecting 
above dactylar articulation as small, acute tooth; 
opposite margin of dactylar articulation bearing 
similar but smaller tooth; superior and internal 
surface of palm papillose with papillae bearing 
heavy but not long setae. Fingers heavy, with 
dactylus slightly curved and fixed finger bearing 
cutting flange on internal edge; tips hooked, 
pointed, and crossing. 
Carpal articles of second leg with ratio: 
10:4.1:3.0:2.6:9.9. 
Third leg with ischium bearing heavy mov- 
able spine; merus 2.9 times as long as broad, 
inferior margins bearing a series of setiferous 
notches and strong terminal tooth; carpus 0.56 
as long as merus, with superior margin bearing 
setae and both superior and inferior margins 
projecting distally into acute teeth; propodus 
0.75 as long as merus, bearing setae on superior 
margin and five pairs of spines on inferior 
margin and two on terminal margin; dactylus 
curved, 0.27 as long as merus, biunguiculate 
with lower unguis small. 
Outer uropod with strong lateral spine; inner 
uropod with a series of strong movable spines. 
Telson 2.7 times as long as broad posteriorly, 
1.5 times as broad anteriorly as posteriorly; 
lateral margins almost straight, posterior mar- 
gin shallowly arcuate; dorsal and terminal spi- 
nules of moderate size. 
DISCUSSION: This species belongs to the 
Macrochirus group, and within the group it is 
obviously most closely related to A. villosus 
(Olivier). Like A. villosus this species has hairs 
over the body surface, a carinate carapace, orbi- 
tal teeth, a strong spine on the basicerite, 
grooves and teeth on the large chela, an acute 
tooth on the merus, and a biunguiculate dactylus 
on the third legs, etc. 
Without a full description of A. villosus, 
point for point comparisons are difficult. The 
differences between the illustrations of Coutiere 
( 1899, seriatim) , taken from Olivier’s type, and 
Bate’s drawings of Paralpheus diversimanus 
(Olivier) (Chalk Rept., pi. 102, fig. 1), which 
Coutiere states is identical with A. villosus 
(1898^:204), are especially confusing. With- 
out any actual specimens of A. villosus for com- 
parison, we have presumed the drawings and 
fragmentary description of Coutiere to be more 
accurate and have used them as much as pos- 
sible in Table 2. 
There is evidently a difference in the large 
chela, especially around the articulation of the 
dactylus, but Coutiere’s sole figure (1899:220, 
fig. 266) is difficult to interpret. 
