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PACIFIC SCIENCE, Vol. XX, April 1966 
figures of Dana lay in the large chela, which he 
shows as having the terminal projection of the 
superior margin at right angles to, but not 
overhanging, the transverse groove. 
Of our efforts to establish a neotype only the 
description above, and the notes following, 
remain. Most of the specimens of the neotypic 
series and the figures of the neotype were 
destroyed. Of the five specimens from Tonga 
saved from the fire, four were incomplete and 
the fifth was too much smaller than Dana’s 
original to be designated as a neotype. This 
last specimen, taken from BT 14, with the 
carapace length of 10 mm, we have drawn and 
are offering to future workers as an example 
of a small specimen from the type locality. 
Alpheus strenuus is a very widespread and 
common species, and was represented in our 
study collection by over 150 specimens. At 
Eniwetok in the Marshall Islands a large number 
were collected from a single environment so 
that the extent of variation could be determined. 
A selection of both sexes and of varying sizes, 
together with random samples from other 
archipelagoes, was carefully measured; all other 
specimens were examined but not measured. 
In all, 20 specimens were measured. The 
form and size of the rostrum and orbital hoods 
were found to be constant. The ratio of the 
antennular articles varied as follows (consider- 
ing the length of the first article to be unity ) : 
1: ( 1.4-2. 1 ) : (0.6-1.0) ; the second article was 
from 1.5 to 2.7 times as long as broad. In 
the specimen described above the second ar- 
ticle was 1.6 times the length of the first and 
2.3 times as long as broad. The tip of the 
stylocerite in all cases reached to, and in a 
few cases slightly beyond, the end of the first 
antennular article. The relative lengths of the 
carpocerite and scaphocerite varied only slightly, 
with the two subequal and slightly longer than 
the antennular peduncle. The spine of the 
basicerite was always present and acute. On 
the large chela the general form remained 
constant, with the upper margin terminating 
in an obtuse tooth that overhung the trans- 
verse groove. The length of the entire chela 
varied from 2.0 to 2.8 times the length of the 
fingers. The merus of the cheliped showed 
greater variation, ranging from 1.7 to 2.8 times 
as long as broad (2.4 in specimen described), 
the larger specimens in general having the 
broader merus. The tooth on the merus was 
of constant development. On the small chela 
all individuals, regardless of size or sex, had the 
balaeniceps- shaped dactylus; however, there 
was variation in the sculpturing of the palm, 
with some individuals showing traces of the 
inferior shoulder. The fingers of this chela 
were 0.40-0.47 the length of the entire chela. 
The merus of the small cheliped, like the merus 
of the large cheliped, was more slender in 
smaller specimens, ranging from 1.8 to 3.0 
times as long as broad; in one specimen the 
tooth on the inferior internal margin was 
obtuse rather than acute. The carpal articles 
of the second legs showed only minor varia- 
tion, with the range of the ratios as follows: 
10: (8-11): (3-4): (3-4): (4-6). On the third 
leg the ischial spine was always present; the 
merus varied from 3-3 to 5.0 times as long 
as broad; the carpus was from 0.5 to 0.8 times 
the length of the merus, and the propodus 0.6 
to 0.7 the length of the same article. The 
maximum breadth of the telson was from 1.8 
to 2.5 times the breadth of the tip. 
The smallest specimen in the collection had 
a carapace length of 3 mm, and was collected 
with the large number of large A. strenuus at 
Eniwetok; this showed the characteristics of 
large specimens except for minor differences 
in the chela. The sculpturing on the large chela 
was not as pronounced as in mature specimens, 
and the termination of the upper margin did 
not overhang the groove; the small chela was 
heavier in proportion to the length, and the 
sculpturing was poorly developed. 
Two varieties or subspecies of this species 
have been described: A. strenuus angulatus 
Coutiere (1905:914) and A. strenuus gala- 
pagensis Sivertsen (1934:3). Coutiere’s sub- 
species was based on three characteristics: first, 
a more slender merus of the third legs, being 
almost 5 times as long as broad; second, a 
more elongate second article of the carpus of 
the second legs, being 1.1 times the length of 
the first; and, third, an inferior margin of the 
merus of the third leg being "plus rectiligne . . . 
