New Monogenetic Trematodes, II — Yamaguti 
423 
long, arcuate, strongly muscular, swollen at its 
forwardly directed base, situated longitudinally 
between right intestinal limb and uterus, con- 
taining prostatic duct and vas deferens, both of 
which are straight, narrow, and close together, 
though convoluted at the anterior base of the 
cirrus pouch. Cirrus spicular, long, enclosed in 
sheathlike genital atrium which extends from 
the posterior distal end of the cirrus pouch to 
the genital pore along the dorsal side of the 
cirrus pouch. This overlapping portion is shown 
separately in the figure. Prostatic reservoir 
elongate, 80-90 X 100-1 30p, situated imme- 
diately in front of base of cirrus pouch. Genital 
pore common, just posterosinistral to pharynx. 
Ovary subglobular, 0.17-0.27 X 0.17-0.22 
mm, situated between two testes and vitelline 
reservoir. Shell gland complex strongly devel- 
oped. Uterus distended with eggs. Eggs rounded 
pyramidal, with triangular base in balsam 
mounts, 0.12-0.14 X 0.12-0.15 mm in life, 
with very fine, long filament at one basal cor- 
ner. Vagina opening ventrally in left submedian 
line a little posterior to intestinal bifurcation; 
vaginal duct thick-walled, running straight 
backward and opening into oval seminal recep- 
tacle which measures 100p by 80p in the type. 
Vitellaria coextensive with intestinal limbs and 
their branches; vitelline reservoir transversely 
elongated, not forming a compact mass, 6 0-90 p 
anteroposteriorly. Paired excretory vesicles con- 
spicuous at level of uterus. 
discussion/ The differences between this 
species and the type species are given in the 
discussion of the type species. 
Pseudallobenedenia n. gen. 
GENERIC DIAGNOSIS: Capsalidae, Benedenii- 
nae. Opisthohaptor discoid, not septate, with 
scalloped marginal membrane, armed with three 
pairs of anchors. Prohaptors not enclosed in 
hoodlike frontal plate. Pharynx muscular, lobed 
anteriorly; intestinal limbs branched, not con- 
fluent posteriorly, though their posterior inner 
branches may anastomose. Testes juxtaposed; 
vas deferens convoluted medial to left intestinal 
limb. Prostatic cells and reservoir outside cirrus 
pouch. Cirrus pouch long ; cirrus extremely 
long, enclosed in extremely long, tubular, genital 
atrium. Genital pore common, sublateral or sub- 
median, level with posterior end of pharynx or 
farther behind. Ovary immediately pretesticular. 
Eggs rounded conical or pyramidal, with very 
long filament at base or basal corner. Vitellaria 
extensive; vitelline reservoir inconspicuous. Va- 
gina opening submedianly ventral to uterus at 
postbifurcal level; receptaculum seminis vagi- 
nae well developed. Gill parasites of marine 
teleosts. 
type species: P. apharei n. sp., on Aphar- 
eus rutilans; Hawaii. 
other species: P. opakapaka n. sp., on Pris- 
tipomoides microlepis; Hawaii. 
3. Lagenivaginopseudobenedenia etelis 
n. g., n. sp. 
Fig. 3 
habitat: Gills of Etelis car bun cuius (local 
name "onaga”) ; Hawaii. 
holotype : U. S. Nat. Mus. Helm. Coll., 
S. Y. No. 47. 
description (based on three whole mounts) : 
Body elliptical, flat, 2. 9-4. 3 mm in total length, 
1.8-2. 3 mm wide in midregion; head and neck 
region contracted and well separated from 
trunk. Opisthohaptor elliptical, 0.53-0.75 mm 
long, with finely scalloped marginal membrane 
30-70p wide and three pairs of anchors; num- 
ber of marginal hooklets not determined; ante- 
rior anchor 0.13-0.17 mm long, sharp-pointed 
anteriorly, nonalate, with two small blunt 
stumpy processes of unequal length at posterior 
end; middle and posterior anchors definitely 
smaller than anterior; middle one slender, 50- 
80p long, with undulating root and minute 
terminal claw; posterior one enlarged and flat- 
tened from side to side basally, 30-40p long, 
terminating in a minute claw. Prohaptors saucer- 
shaped, paired, close together, 0.19-0.23 X 
0.36-0.42 mm, containing glandular tissue 
which extends not only backward but also in- 
ward to be confluent in the median line pos- 
terior to somewhat incised frontal margin. Two 
pairs of eyespots present anterodorsal to phar- 
ynx. Pharynx spherical, 0.27-0.29 X 0.3-0.31 
mm, glandular rather than muscular, papillate 
internally. Esophagus practically absent ; ceca 
with numerous dendritic outer branches, not 
confluent posteriorly. 
