Flora of Josephstaal, Papua New Guinea 
Lae National Herbarium 
1 3633 Thelypteridaceae 
Plesioneuron tuberculatum (Cesati) Holttum 
keys to genus and the species on Holttum 
Madang Province, Josephstaal FMA area, near 
Kumamdeber, along streambed flowing to SW of 
expedition Camp 1 and returning along ridgeline; 
streambed bordered by many landslides, mostly with 
early successional vegetation but also with mature 
forest patches on higher slopes; Kumamdeber GPS 
281407 E, 9504560 N, elev. 530 ft. (ca. 160 m), the 
stream joins a larger tributary at GPS 28151 E, 
950298 N. 
coll, sequence 13608-13646; terrestrial fern on 
landslide colluvium; rhizome close-creeping; stipes 
green or atropurpureous, unarmed, stele U-shaped; 
leaves herbaceous, opaque green both surfaces, 
pinnae numerous and stalked; venation pinnatiform, 
free to the margins; sori costal, indusiate, without red 
glands, sporangial body with globose sessile glands. 
coll.: W. Takeuchi, J. Wiakabu, M. Gorrez, A. Towati 13633 
date: July 31, 1999 
det.: WT; Jan. 4, 2000 
Flora of Josephstaal, Papua New Guinea 
Lae National Herbarium 
1 3634 Moraceae 
Ficus botryocarpa Miq. 
var. subalbidoramea (Elmer) Corner 
Madang Province, Josephstaal FMA area, near 
Kumamdeber, along streambed flowing to SW of 
expedition Camp 1 and returning along ridgeline; 
streambed bordered by many landslides, mostly with 
early successional vegetation but also with mature 
forest patches on higher slopes; Kumamdeber GPS 
281407 E, 9504560 N, elev. 530 ft. (ca. 160 m), the 
stream joins a larger tributary at GPS 28151 E, 
950298 N. 
coll, sequence 13608-13646; arborescent; leaves 
papery, adaxially opaque very dark green, abaxially 
light green; figs from older wood, summit depressed, 
receptacle opaque dark green with lighter colored 
lenticels. 
coll.: W. Takeuchi, J. Wiakabu, M. Gorrez, A. Towati 13634 
date: July 31, 1999 
det.: WT; Dec. 13, 1999 
