Flora of Josephstaal, Papua New Guinea 
Lae National Herbarium 
1 3953 Pteridaceae 
Pteris ligulata Gaud. 
Madang Province, Josephstaal FMA area, colls around 
expedition Camp 2 at the Guam River; centered on 
GPS 280100 E, 9497596 N, elevation ca. 50-100 m; 
generally from lower slopes of natural-growth foothill 
forest. 
coll sequence 13953-14003; terrestrial fern on 
riverbank; stipes atropurpureous; leaves herbaceous, 
upper surface opaque medium green, yellow-green 
beneath. 
coll: W. Takeuchi, J. Wiakabu, M. Gorrez, A. Towati 13953 
date: Aug. 13, 1999 
det.: WT; Jan. 2000 
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