Flora of Josephstaal, Papua New Guinea 
Lae National Herbarium 
1 3959 Leeaceae 
Leea indica (Burm. f.) Merr. 
Madang Province, Josephstaal FMA area, colls around 
expedition Camp 2 at the Guam River; centered on 
GPS 280100 E, 9497596 N, elevation ca. 50-100 m; 
generally from lower slopes of natural-growth foothill 
forest. 
coll sequence 13953-14003; small shrub; leaves newly 
flushed, adaxially pale green, abaxially brownish-green, 
rachis at least 2-jugate; flowers yellow-green in bud, 5- 
merous. 
coll: W. Takeuchi, J. Wiakabu, M. Gorrez, A. Towati 13959 
date: Aug. 13, 1999 
det.: WT; Dec. 20, 1999 
Flora of Josephstaal, Papua New Guinea 
Lae National Herbarium 
13960 Euphorbiaceae 
Macaranga quadriglandulosa Warburg 
var. quadriglandulosa 
keys to this result on Whitmore 
Madang Province, Josephstaal FMA area, colls around 
expedition Camp 2 at the Guam River; centered on 
GPS 280100 E, 9497596 N, elevation ca. 50-100 m; 
generally from lower slopes of natural-growth foothill 
forest. 
coll sequence 13953-14003; subarborescent in 
regrowth; branchlets green, smooth, angular when 
dried; leaves papyraceous, adaxially opaque medium 
green, abaxially glaucescent, yellow-pulverulent; fruits 
green, armature of flexible and squamate spines; bracts 
fleshy, yellow-green. 
coll: W. Takeuchi, J. Wiakabu, M. Gorrez, A. Towati 13960 
date: Aug. 13, 1999 
det.: WT; Dec. 10, 1999 
