6 Lawson . — The Gametophytes , Fertilization and 
and the cells near the tubes become thoroughly disorganized. The number 
of tubes varied from one to four, but more generally only one or two were 
found in a single nucellus. As the tube advances, the body-cell and the 
tube and stalk-nuclei were invariably situated near the tip, as shown in 
Fig. 5 . When the tip of the tube has penetrated through the nucellar 
tissue and has reached the space between the nucellus and the archegonial 
chambers, it becomes quite distended, as if a great osmotic pressure were 
acting from within. It would seem that the sudden release from the con- 
finement of the surrounding nucellar tissue permitted the wall of the tip of 
the tube to stretch, allowing the latter to take on an almost spherical or 
bulb-shaped form, as represented in Figs. 5 and 6 . It will also be seen 
from these figures that the bulk of the cytoplasm is at the tip of the tube, 
with a thin layer lining the wall. 
As soon as the tube reaches this condition the nucleus of the body-cell 
enlarges and prepares for division. This stage is shown in Fig. 6 . I was 
unable to find the spindle of the dividing body-nucleus, but a careful study 
of the cytoplasm before and after the spindle stage revealed nothing that 
would indicate the presence of blepharoplasts, which occur during the 
division of the body-cell in the Cycads (Ikeno, ’98 ; Webber, 97) and 
Ginkgo (Hirase, ’95). The cytoplasm immediately before division was 
finely granular at the periphery, but distinctly fibrilar in its structure in 
the vicinity of the nucleus, which indicated the preliminary stages in the 
formation of the spindle. 
The result of the division of the body-nucleus proved very interesting 
because it differed in a marked degree from that reported for Taxus 
(Belajeff, ’93) and Torreya taxifolia (Coulter and Land, ’05). It will be 
remembered that in Taxus this division results in the organization of two 
male cells of unequal size. In Torreya taxifolia a similar condition 
prevails, although Miss Robertson (’04) reports an equal division of the 
body nucleus in Torreya californica , and thus similar to the conditions 
found in Cephalotaxus Fortunei by Arnoldi (’00). Miss Robertson, however, 
believes that only one of the male nuclei enters the archegonium. The 
discovery of Juel (’04) that the body-cell in Cupressus Goweniana gives rise 
to a complex of cells is an interesting and important one, especially as 
Thomson (’05) and Lopriore (’05) recently report that in Agathis australis 
and in Araucaria the pollen-tube may contain six or seven nuclei. In these 
latter cases it will be interesting to learn whether these nuclei are vegetative 
or generative. 
In Cephalotaxus drupacea I was able to follow a very complete series 
of stages after the division of the body-cell nucleus, and my observations 
are these : — There is no cell-plate formed between the two male nuclei, 
and consequently there are not two male cells formed, but simply two large 
nuclei, which lie within the old membrane of the body-cell. These nuclei 
