352 Fraser . — On the Sexuality and Development of 
Vegetative Mycelium. 
The cells of the mycelium are multinucleate (Fig. 3) and show 
numerous conspicuous granules, often aggregated on the transverse walls, 
and corresponding, no doubt, to those observed in several other members 
of the Pezizineae. No organs of reproduction other than the ascocarps 
were observed in this form. 
Development of the Apothecium. 
The archicarp 1 arises as a side branch from a vegetative hypha 
(Fig. 4), and becomes divided into two, then three (Fig. 5), and finally five 
or more cells. The terminal cell or ascogonium is oval in shape and larger 
than the others, and, as development proceeds, it becomes filled with dense 
contents. In cut and stained preparations it is seen to contain numerous 
nuclei ; the cytoplasm is finely granular and shows a number of small, 
rounded vacuoles (Fig. 6). 
Hyphae grow out from the lower cells of the archicarp and from the 
branch which bears it, and form a dense weft, above which the ascogonium 
rises. 
So far the archicarp resembles that described by Woronin (22) for 
Lachnea scutellata , Gill. (= Peziza scutellata , L.), where a large, oval asco- 
gonium, with granular, somewhat vacuolate contents, terminates the usually 
three-celled archicarp. Branches, one of which Woronin thinks may be 
the antheridium, grow up from the lower cells and cover in the, as yet, 
unbranched ascogonium. Owing to the density of the investing hyphae 
further development could not be traced. 
In Lachnea ster corea , however, the ascogonium, before it is surrounded 
by hyphae, itself sends out a stout branch (Fig. 7), into which several 
nuclei pass (Fig. 8). This branch is cut off from the ascogonium by a wall 
on which granules may often be observed. It grows in length and becomes 
divided, first into two nearly equal cells, and eventually into four, five, or 
six cells (Fig. 9). Development probably takes place with some rapidity, 
for, though the unicellular stage was often found, the stage with two cells 
was only twice observed. 
When fully developed this branch consists, as a rule, of five or six 
cells ; those towards the archicarp are narrow, and, though always multi- 
nucleate, contain few nuclei. The terminal cell is considerably larger and 
its nuclei are much more numerous (Figs. 9, 10). In the early, unicellular 
stage the nuclei of the branch are quite similar to those of the ascogonium, 
1 This term is used for the whole fertile branch other than the antheridium, while the term 
ascogonium is restricted to the reproductive cell or cells containing female nuclei (Blackman and 
Fraser (4)). 
