Campbell. — Studies on some Javanese Anthocerotaceae. I. 469 
third section of the genus, and, together with the other characters, would 
seem to constitute sufficient ground for separating these species as a distinct 
genus, as Leitgeb (loc. cit. p. 27) believed should be done. The discovery 
of the multiple chromatophores is important, as it breaks down one of 
the barriers separating the Anthocerotaceae from the other Archegoniates ; 
but we still believe that the differences are sufficient to warrant the retention 
of the class Anthocerotes. 
Megaceros, a new genus of Anthocerotaceae. 
Shortly after the writer’s arrival in Java, in March, 1906, while on 
a botanizing excursion to the Tjiapus Gorge at the base of Mount Salak, 
near Buitenzorg, a number of species of Anthoceros were collected, and 
among them was one growing among mosses upon a fallen log. As this 
was a position very similar to that in which the West Indian specimens were 
collected, it was hoped that this might prove to be a similar form, and 
an examination after returning to Buitenzorg showed that such was the 
case. These specimens were fruiting abundantly, and it was at once 
evident that the plant belonged to the section with spiral elaters and no 
stomata, thus approaching Dendroceros. Later collections at Tjibodas 
showed that a similar species was not rare in the neighbourhood, and 
plants were repeatedly collected at various points from the immediate 
neighbourhood of the laboratory (altitude about 1,450 metres) to a short 
distance below Kandang Badak, which is about 2,500 metres high. A 
comparison of these specimens with those collected on the Salak showed 
that they were evidently specifically distinct, although much resembling 
the species from the lower level. 
The Tjibodas material showed a good deal of variation as to habitat. 
Most commonly, as in the specimens first collected, the plants grew on 
rotten logs ; but sometimes they were found upon the earth, and, in a few 
instances, upon boulders. The latter specimens had a thicker thallus than 
the others, but otherwise were not perceptibly different, and probably 
are not specifically distinct. A number of specimens with quite different 
chromatophores were found in the material when it was examined upon 
the writer’s return to America, but these were sterile, and so could not 
be compared with the others. It is possible that they represent a third 
species, but at present this can only be conjectured. 
The multiple chromatophores, together with the Dendroceros - like 
type of sporogonium, i. e. spiral elaters, green spores, and the absence 
of stomata, seems to warrant a separation of the group from the other 
species of Anthoceros , as Leitgeb suggested should be done. The name 
Megaceros is here proposed for all the species of this group. The name is 
selected on account of the very large size of the sporophyte in the commonest 
Javanese species. This in some instances reaches a length of 9 cm. and 
