Campbell. — Studies on some Javanese A nthocerotaceae. I. 471 
M. Tjibodensis. In both species the usual Nostoc colonies occur, but 
these are not especially noticeable. 
The antheridia are produced singly in the usual chamber, and are 
large enough to be seen with the naked eye, in some cases having a 
diameter of about 170 fi. Stephani states that in M. Stahlii they do 
not occur upon the same branches of the thallus as the archegonia, but 
in all forms studied by the writer they occurred upon the saAe shoot. 
The plants are markedly proterogynous, however, unlike most species 
of Anthoceros, where the antheridia are formed first. It may have been 
the overlooking of this fact that led Stephani to suppose that they were 
borne upon special branches. 
The Chromatophores. 
A careful study of the chromatophores of both of the Javanese species 
was made. These are most different from the ordinary Anthoceros type 
in M. Tjibodensis. In most species of Anthoceros , as is well known, each 
cell contains a single large flat chromatophore, in the centre of which is 
a very distinct pyrenoid, the whole closely resembling the chromatophores 
of many green Algae, especially many Confervoideae, like Stigeocloitium 
or Coleochaete. A careful study of M. Tjibodensis shows that not only 
is the single chromatophore replaced by several, sometimes as many as 
twelve, but all trace of a pyrenoid has disappeared, so that the chromato- 
phore is in all respects like that found in the other Archegoniates. The 
pyrenoid also seemed to be absent from the chromatophore of M. Vin- 
centianus and M. jlavens , of which dried specimens were examined ; but 
in M. Salakensis (Figs. 13 to 17) there was an aggregation of small starch 
granules about a central area in the larger chromatophores, which had the 
appearance of a pyrenoid, although it was much less definite than is usually 
the case in most species of Anthoceros. 
M. Tjibodensis , although in a few of the superficial cells a single 
chromatophore may occur, in most cases has from two to four, while 
the number is much greater in the inner cells of the thallus. In the 
superficial cells of the upper side (Fig. 9) there are most commonly two 
chromatophores. Sometimes there is a single one and sometimes three. 
In the superficial cells of the ventral side the number ranges from two 
to four (Fig. 10), and they are smaller, but otherwise resemble those of 
the dorsal surface. They are in both cases thin flat plates, showing a 
uniform finely granular appearance, but no trace of anything like a pyrenoid, 
nor were large starch granules observed. In the cells immediately below 
the surface there are usually four smaller chromatophores, while in the 
large inner cells (Fig. 11) they are much more numerous, but of smaller 
size, and often quite round, closely resembling the typical chlorophyll 
bodies of higher plants. The nucleus (n) in all the cells is very small, 
