Campbell. — Studies on some Javanese Anthocerotaceae. /. 481 
(Fig. 63 a). In the next stage (b) the chromatophore divides into two, 
which assume an oval form, and show clearly a number of starch granules 
imbedded in them. The two chromatophores are in Megaceros connected 
by a strand of fine fibrils, which Davis does not seem to have seen in 
Anthoceros. About this time there is a marked appearance of synapsis 
in the nucleus (Fig. 63 b), and Davis says that this is a constant phenomenon, 
and regards it as entirely normal. The two chromatophores divide again 
and assume a position marking the tetrad arrangement of the future spores. 
They are connected, as in the case of the two chromatophores, by strands of 
fibrils, and have very much the effect of the four nuclei connected by fibrils 
which are seen in the ordinary type of spore division. The nucleus now 
divides, as Davis has shown, by successive mitoses, with a resting-stage 
between, and the four nuclei arrange themselves at the four points occupied 
by the chromatophores, after which the cell-wall separating the four spores 
of the tetrad are developed. It was noted that before the division began 
there was already a slight lobing of the cell, such as is so common in the 
Jungermanniales, marking the position of the spores. This has also been 
observed by the writer in Targionia. 
The outer membrane of the spore becomes somewhat thickened, but 
remains thinner than is the case in Anthoceros and Notothylas. In this 
respect Megaceros approaches Dendroceros , and the spores agree with those 
of Dendroceros also in the development of chlorophyll, which is wanting in 
the ripe spores of all typical species of A nthoceros. 
The elaters, as in Dendroceros , are as a rule composed of several cells, 
and are simply larger or smaller fragments of the net of sterile cells which 
surrounds the spores. This becomes especially clear in M. Salakensis , 
where the elates are usually branched (Fig. 72). In M. Tjibodensis they 
are rather more slender and seldom show any branches. The characteristic 
thickened spiral upon the walls of the elaters first becomes evident about 
the time that the division of the spores is complete. In both species the 
ripe spores show fine spinules or papillae upon the outer surface, these 
being more numerous, finer, and more regular in M. Salakensis. The 
superficial cells of the sporogonium develop thick walls and assume 
a brownish colour as the sporogonium ripens, but no trace of the stomata 
characteristic of the typical Anthoceros sporogonium is developed. The 
two species examined differ in the form of the superficial cells of the 
sporogonium, which are about twice as long in M. Tjibodensis as in 
M. Salakensis. In the former species there are two to six chromatophores 
in each cell of the assimilative tissue of the sporophyte, the commonest 
number being four. 
As the sporophyte develops, rhizoids are formed in great numbers 
from the lower surface of the thallus below the foot of the sporophyte, the 
growth of which in M. Tjibodensis causes a marked swelling upon the lower 
