Campbell. — Studies on some Javanese Anthocerotaceae. I. 483 
origin of the sporogenous tissue it is more like Dendroceros , with which it 
agrees also in the green spores, the spiral elaters, and in the absence of 
stomata. The great development of the sporogenous tissue, as well as the 
large amount of sporogenous tissue developed in the apical region of the 
sporophyte, are more like Nototkylas. 
Summary. 
1. Probably all of the species of Anthoceros in which stomata are 
absent from the sporogonium, and which have spiral elaters, should be 
separated as a new genus. The name Megaceros is proposed for this. 
1. Two species from Java were carefully studied ; these appear to be 
undescribed, and the names M. Tjibodensis and M. Salakensis are pro- 
posed for them ; the former is probably closely allied to M. Stahlii (Steph.), 
but seems to be distinct. 
3. Both species show multiple chromatophores, and this peculiarity is 
probably common to all members of the genus. Two tropical American 
species, M. Vincentiamis and M. Jlavens , show the same character, and it 
is also known in M. giganteus. In M. Tjibodensis as many as twelve 
chromatophores may be found in a cell. Pyrenoids were absent in all of 
the forms studied except M. Salakensis where there was some evidence that 
a pyrenoid was present. 
4. The form of the thallus and the apical growth are like those of the 
typical Anthoceros. 
5. Both of the Javanese species are monoecious, but strongly protero- 
gynous ; the archegonia resemble those of Anthoceros , but the large solitary 
antheridium is more like that of Dendroceros. 
6. The early divisions in the embryo and young sporophyte most 
nearly resemble Dendroceros ; the extent of the sporogenous tissue, however, 
is much greater, and most like Notothylas. 
7 . There is no evident relation between the early divisions of the 
sporogenous tissue and the distribution of the fertile and sterile cells. The 
latter form an irregular net enclosing the spore-mother-cells. 
8. As in Anthoceros the primary chromatophore of the spore-mother- 
cell divides into four before the nucleus divides ; there is a slight lobing 
of the mother-cell before division takes place ; the spores are small, thin- 
walled, with fine papillae or spinules upon their surface, and contain a large 
chloroplast. 
9. The elaters are multicellular, and closely resemble those of 
Dendroceros ; they are branched in M. Salakensis , but unbranched in 
M. Tjibodensis . 
10. There is a large amount of green tissue developed in the sporophyte, 
but no stomata are present ; in M. Tjibodensis the cells of the sporophyte 
