484 Campbell ’■ — Studies on some Javanese Anthocerotaceae. L 
contain 2-6 chromatophores ; the dehiscence of the capsule is either by two 
valves (M. Tjibodensis), or along one suture (M. Salakensis). 
11. The foot is very large, and the surface cells are developed into 
extensively branched rhizoid-like outgrowths ; rhizoids develop in great 
numbers from the lower surface of the thallus immediately below the foot. 
Megaceros , gen. nov. 
Thallus large in all of the Javanese species, closely resembling 
Anthoceros , both in form and in the apical growth ; chromatophores 
several (2-12) in the inner cells, and usually more than one in the super- 
ficial ones ; pyrenoids are usually absent. Plants (always ?) monoecious ; 
antheridia large, solitary ; sporophyte large, epidermis without stomata ; 
spores small, thin-walled, papillate or echinulate, at maturity containing 
a single large chloroplast ; elaters either branched or unbranched, multi- 
cellular, with a distinct spiral band like that of Dendroceros. 
Probably all of the species of Anthoceros in Gottsche’s third section of 
the genus should be transferred to Megaceros. 
M. Tjibodensis, sp. nov. 
Thallus 2-5 centimetres long ; ecostate, 8-12 cells thick in the middle. 
Margin strongly lobed, and laciniately fringed ; monoecious, but strongly 
proterogynous ; antheridia nearly globular, upright, about 135 n in diameter, 
short stalked; involucre 10-15 mm. long; sporophyte very long, 6-9 c. ; 
spores 28/1, somewhat irregular in shape, papillate: elaters long -and 
slender, sometimes 300 [x in length, unbranched ; epidermal cells of sporo- 
phyte long and narrow. Growing on logs, upon the ground, and upon 
boulders. Tjibodas ; also below Kandang Badak on the slopes of Mount 
Gedeh, Java; April, May, 1906. 
Probably near to M. (Anthoceros) Stahlii (Steph.), but differs in the 
larger size and greater thickness, the larger spores, and much longer 
elaters. 
M. Salakensis , sp. nov. 
Thallus smaller than M. Tjibodensis , about 2-3 centimetres long, 
8-10 cells thick in the middle, crenately lobed but not laciniate ; chromato- 
phores 1-2 in the superficial cells, 2-6 in the inner ones, a pyrenoid probably 
present in the larger ones ; monoecious, proterogynous ; antheridia oval, 
about 180/x in length exclusive of the curved pedicel which is much longer 
than in M. Tjibodensis , and attached at the forward part of the antheridial 
chamber ; sporophyte much shorter than in M. Tjibodensis , 3-4 centimetres 
long ; spores 28 [x, nearly round, and finely echinulate ; elaters branched, 
somewhat thicker than in M. Tjibodensis , epidermal cells of the sporophyte 
about half as long as in M. Tjibodensis. On a dead log, among mosses ; 
Tjiapus Gorge, foot of Mount Salak, Java ; March, 1906. 
