222 
thick-walled, and only slightly larger than the 
oral pouches, 24 X 40 p, the mean diagonals 
of the pouches being as much as 70% that 
of the pharynx. The oesophagus is 0.1 6 mm 
long and unbranched, and bifurcates into the 
intestinal crura immediately behind the male 
genital pore. At the posterior end of the neck 
region, the crura have numerous complex outer 
branches and a few simple inner branches ; they 
are particularly extensive on the side bearing the 
clamp row. The crural ends are close together 
near the tip of the body, but they are not con- 
fluent across the median line, nor were any 
intercrural bridges seen. 
The haptor, represented by the unilateral 
clamp-bearing flange on the hindbody and in- 
cluding the lappet, is slightly less than half the 
total length of the slightly contracted worm 
(Fig. 18), but in the extended condition it is 
only 36%. The long clamp row (usually on the 
left) is 0.825-1.35 mm long, bearing 27-35 
almost sessile clamps. The inhibited side of the 
haptor is represented in all specimens by 3 rem- 
nant clamps, and together they make a row only 
0.10-0.12 mm long. Occasionally (as shown in 
Fig. 18), there may appear to be only 2 rem- 
nant clamps in the short row (usually the right 
side in my collections), but in Figure 20 it will 
be seen that the first two primary clamps are 
relatively minute and lie close together near the 
median line between the second pair of primary 
clamps — virtually the end clamp of each row. 
In most specimens the three clamps of the in- 
hibited side are subequal and in a linear series, 
as in Figure 19. 
The terminal lappet is short, narrow, and 
cylindrical, 0.82 X 0.25 mm-0.98 X 0.48 mm, 
armed with two pairs of symmetrically arranged 
anchors. The anterior anchors have their hooks 
only slightly sickle-shaped (one-quarter to one- 
third of a circle), with a knoblike spur at the 
top of the slender handle, which may be less 
than one-half the total length of the anchors 
(40 p). The posterior anchors are almost C- 
shaped, with a reflexed strongly rounded hook 
but a very short handle (total length 12-16 p). 
Both pairs of anchors are similar to those fig- 
ured by Tripathi (1959), but while the anterior 
pair is like those of Engrauliphila the posterior 
pair resembles only those of En t granite ola micro- 
pharyngella. 
PACIFIC SCIENCE, Vol. XXI, April 1967 
The clamps in the long row are 44 X 52 p- 
35 X 75 p, and in the short row 20 X 28 
30 X 48 p. Thus, the primary clamps of both 
rows are smaller and only slightly wider than 
long, but those in the long row are graded as 
usual, the largest being on either side of the 
middle region, and more than twice as wide as 
long. The braces are situated across this longer 
diameter; they are nearly straight with bent ends 
lacking the bent articular facet, though they do 
meet medially near the level of the divergent 
V-like arms of the dorsal appendix. The ventral 
arm of the spring is very slender, with or with- 
out a very slight enlargement at its end, but this 
extremity is always with a minute notch, never 
truly bifurcated (see Figs. 21 and 22). 
There are 9-12 irregularly oval testes, 60 X 
75 p-75 X 150 p, arranged in two files and 
not in a single mass as depicted for the geno- 
type. The row on the side opposite the clamps 
is completely postovarian, while that nearer the 
clamp row extends forward to the middle of the 
ovary along the outer edge of the median vitel- 
line duct; these parovarian follicles were not 
found in E. thris socles. The vas deferens orig- 
inates from the anteriormost testes of the par- 
ovarian file, runs forward parallel to the uterus, 
and opens into a zigzag ejaculatory duct which 
in turn opens into the penis. No vesicula sem- 
inalis was seen. The muscular penis is small 
and conical, without forceps, and opens into the 
circular atrium masculinus, which is surrounded 
by a muscular ring 20-24 p in diameter, sit- 
uated at about 0.16-0.19 mm from the anterior 
end of the body. The penis itself bears around 
its widest diameter a corona of 12 sharp conical 
spines pointing vertically from the ventral sur- 
face; the spines are nearly straight. Figures 23 
and 24, drawn from ventral and dorsal aspects, 
are intended to demonstrate the entire absence 
of forceps on the penis head. 
The ovary is in the form of an elongated in- 
verted U, with the distal (inner) arm wider 
and containing a number of large ova, and the 
proximal end slightly swollen and overlapped 
by the anterior testes of the postovarian file. 
The short and narrow oviduct expands slightly 
to form a muscular ovijector, before opening 
into the vitelline ampulla (Fig. 25). The wide 
median uterus arises from the outer margin of 
the fertilization chamber in the ootype, runs 
