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PACIFIC SCIENCE, Vol. XXI, April 1967 
of that of the pharynx. The oesophagus is long, 
narrow, and unbranched, and bifurcates into the 
intestinal crura well behind the male pore. The 
crura have relatively few regular outer branches, 
extend backward to the level of the proximal 
end of the short row of clamps, and terminate 
independently, but close to each other, without 
inflations. The crus adjacent to the longer clamp 
row sends out wide oblique branches to each of 
the anterior clamps. 
The haptor in Pellonicola , in contrast to that 
in all the previous genera, is distinct from the 
body proper and constitutes a posterior tail. Its 
right and left haptoral frills are parallel, but 
one is only half as long as the other: in my 
specimens, the left frill is always the longer, 
0.45-0.93 mm; the right frill range is 0.225- 
0.456 mm and it bears 8-10 clamps, 36 X 
20 p-44 X 36 p, while the left frill has 17-22 
clamps, 36 X 24 P - 48 X32 p. 
The terminal lappet apparently is unique in 
its telescopic arrangement and consists of an 
oblique plaque capable of being retracted as a 
whole into the hollow end of the haptor (Fig. 
28) ; it is 60 X 28 p-68 X 36 p, and is 
armed with the usual two pairs of anchors 
(Figs. 28 and 29). The anterior pair is dis- 
tinctively shaped like a sickle, 40-44 p long, 
with the handle about equal in length to the 
hook, but the spur root is more than half as 
long as the handle and projects at right angles 
from it. The tip of the spur is slightly curved 
but does not actually pivot backward, as in 
Engraulzcola micropharyngella (Fig. 3), which 
otherwise it most resembles in this series (par- 
ticularly on account of its sickle hook, which 
is slightly more than a half circle, although 
the handle is more slender and relatively 
longer). The posterior anchors are bent in a 
deep C -shape and their over-all length is only 
16-20 p with virtually no handle, as in Engrau- 
liscobina triaptella (Fig. 20, compare with Fig. 
29) . The placing of the anchors is invariable; 
the anterior pair is always directed outward and 
the posterior pair inward, their hooks nearly 
touching, in all genera. 
The clamps are, exceptionally, slightly longer 
than wide, but their structure is very different 
in detail from any of those previously discribed : 
the median spring is highly modified, the ven- 
tral arm being vase-shaped in outline and ap- 
parently split longitudinally, with each half 
bowed outward ; the tip is bifurcated. The short 
arm of the median spring, which has radiating 
tendonous striae at its distal end, has an ap- 
pendix which in some specimens is typically 
U-shaped (Fig. 31), but in others is shaped 
like half a Maltese cross (Fig. 30). The braces 
(oblique sclerites) are stout and slightly wavy, 
and, although their inner ends touch in the 
middle line, there is no sharp bend here to 
form the familiar articular facet. The dorsal 
arm of the ventral jaw sclerite is unusually 
small, often appearing as a mere knob at the 
region of articulation of the dorsal and ventral 
jaws (compare Figs. 28 and 30). 
There are 34-52 testes arranged in 2-3 rather 
regular files. On the side of the uninhibited 
clamp row there is a single file of a few testes 
extending parovarially up to the level of the 
transverse vitelline duct on the left side (Fig. 
33) ; the anterior testes are largest (48 X 
68 p), while the posterior ones are smaller 
(16 X 20 The vas deferens is long and 
zigzag, arising from between the anterior testes 
and running forward; in front of the ovary it 
widens into a large vesicular duct to about the 
level of the lateral vaginae, behind the in- 
testinal bifurcation. This part probably func- 
tions as a seminal vesicle. Thence the vas 
deferens continues forward, to open into the 
base of the penis. The muscular cuplike penis 
is armed equatorially with a corona of 10-12 ! 
recurved spines, 10-12 p long (Fig. 32), but 
the penis head within is devoid of forceps. The 
median ventral atrium masculinus has a rim of 
radial muscle fibres and is situated at about 
0.12-0.27 mm from the anterior end of the 
body, in front of the intestinal bifurcation. 
As usual, the ovary is in the form of an in- 
verted U and is situated in the middle third 
of the body, slightly shifted to the right side; 
the proximal region is oval and lies immediately 
in front of the testicular zone on the right side; 
it is long and narrow, and the distal limb is 
short (two-thirds the length of the longer Fmb) 
and contains large ova, each 12-14 p in diam- 
eter. The short and narrow oviduct descends 
from the distal end of the ovary and enlarges 
into a well-demarcated ovijector at about the 
middle of its length and thence continues back- 
ward to open into the median vitelline duct, 
