Systematics of Prickly Sculpin, I — Krejsa 
243 
onymy of C. gulosus') . Lindsey, 1957:657 
(British Columbia: Columbia R. ; Fraser R.; 
Skeena R.; coast drainages south of Skeena; 
Nass R. ; Stikine R. ; Peace R.). Wilimovsky, 
1958:62 (key to species in Alaska; southeast 
Alaska to California). Shapovalov, Dill, and 
Cordone, 1959:173 (listing only). Carl, Clem- 
ens, and Lindsey, 1959:158 (description; 
Pacific drainages from Chilkoot L., Alaska, to 
Ventura R., California. In British Columbia: 
lakes and rivers of the Columbia, Fraser, Dean, 
Skeena, Nass, and Stikine systems; coastal 
rivers of the mainland and Vancouver Island, 
and Queen Charlotte Islands; headwaters of 
Peace R. system from Summit L. to McLeod 
L.). McAllister and Lindsey, 1959:70 (de- 
scription; synonymy; intraspecific variation; 
localities as in Carl, Clemens, and Lindsey, 
1959). McAllister, 1960:42 (collection in salt 
water, Pt. Atkinson, British Columbia). Bond, 
1961:36 (key to species in Oregon; prickling 
variation; Columbia R. drainage). Bond, 1963: 
79 (synonymy; life history observations; oxy- 
gen, temperature, and salinity tolerance of 
adults; fish associates; habitat preference; lists 
35 new collection localities in coastal Oregon, 
30 localities from Columbia R. drainage in 
Oregon, and also some lakes in the southwest 
corner of Rainier National Park, Washington). 
Bailey and Bond, 1963:19 (recognition of 
several species groups within Coitus in western 
N. America; characters and list of species in 
the " asper species group”). Krejsa, 1965:1-109 
(synonymy; nomenclatural history; distribution; 
life history; morphological variation; salinity 
tolerance of eggs; phylogeny of C. asper and 
closely related species). Bohn and Hoar, 1965: 
977 (salinity effects on iodine metabolism; 
physiological divergence of inland and coastal 
C. asper). 
Centridermichthys asper. Richardson, 
1844:76 (River Oregon [= Columbia R.]). 
Gunther, 1860:170 (description; synonymy; 
fresh waters of the Oregon and Washington 
Territories). Lord 1866^:130 (life history; 
spawning behavior, in part; Puget Sound; 
| "streams flowing through the Sumass and 
Chilukweyuk prairies” [Sumas and Chilliwack 
R. ?], British Columbia; in part, all streams 
east and west of the Cascades). 
Cottopsis asper. Girard, 1851^:303 (intro- 
duction of Cottopsis gen. nov. ; synonymy; 
limited to River Oregon [= Columbia R.]). 
Girard, 1851c:185 (not seen). Girard, 1852:61 
(definition of Cottopsis gen. nov., based on 
Richardson’s description of Cottus asper ; syn- 
onymy; Columbia R.). Girard, 1859:51 (de- 
scription; synonymy; based on 8 specimens 
from Astoria and Fort Dalles, Oregon, and 
Fort Steilacoom, Puget Sound, Washington 
Territory). Suckley (in Cooper and Suckley, 
1859) 1859:351, and Suckley, 1860:351 (de- 
scription; synonymy; small fresh-water streams 
emptying into Puget Sound; Ft. Steilacoom; 
and Columbia R., 200 miles above mouth). 
Jordan and Jouy, 1882:5 (Puget Sound; Co- 
lumbia R. ; Mare Island and Sacramento R., 
California) . 
Uranidea asp era. Jordan and Gilbert, 
1883:694 (description; synonymy; streams 
west of the Sierra Nevada and Cascade Moun- 
tains). Jordan, 1885: 110 (subgenus Cottopsis 
and a list of species therein). 
Trachidermus richardsonii. Heckel, 1840:162 
(synonymy; Columbia R.), (not Cottus richard- 
soni of Agassiz, 1850). Note: Girard, 1852:62, 
erred in reporting the date of Heckel’ s paper 
as 1837, and in the spelling of Trachidermus. 
Cottopsis parvus. Girard, 1856^:144 (orig- 
inal description ; Presidio on San Francisco 
Bay, California). Girard, 1857:11 (descrip- 
tion; Presidio, and Monterery, California). 
Girard, 1859:54 (description; synonymy; 
Monterey, Presidio, Fort Reading, and Peta- 
luma, California). Cooper, 1868:492 (listing 
only). Jordan, 1877:5 (as the young of Cottop- 
sis asper). 
Centridermichthys parvus. Gunther, I860: 
170 (description; synonymy; fresh waters of 
California). Lord, 1866^:352 (listing; "fre- 
quenting the same localities as . . .” Centri- 
dermichthys asper). 
Uranidea aspera var. parvus. Jordan and 
Gilbert, 1883:694 (Sacramento R. forms). 
Cottus gulosus parvus. Jordan and Ever- 
mann, 1898:1945, and Jordan, Evermann, and 
Clark, 1930:383 ( Cottopsis parvus’. Monterey, 
Presidio, Fort Reading, and Petaluma, Cali- 
fornia) . 
