CORRIGIOLA 
l 3 
Genus 4. Corrigiola 
Corrigiola L. [Gen. PI. 340 (1737) non Dillenius;] Sp. PI. 271 (1753) et Gen. PI. ed. 5, 132 (1754); Pax 
in Engler und Prantl PJlanzenfam. iii, pt. 1 b, 88 et 90 (1889); Rouy FI. France x, 11 (1910). 
Annual or perennial herbs. Leaves all or mostly alternate, glabrous ; stipules scarious ; petioles 
short or none. Laminae linear or oblong, rather -succulent. Sepals 5, persistent, connate at the 
base, obtuse, margins white. Petals 5, oblong, white. Stamens 5. Style short or absent. Stigmas 3, 
very short. Fruit somewhat trigonous, indehiscent, 1 -seeded. Pericarp thick, rather rugose. Seed 
large, 3-angled, suspended by a basal funicle. Testa membranous. Embryo annular. 
About 12 species; central and southern Europe; South America; southern Africa. Only 
British species : — C. littoralis. 
I. CORRIGIOLA LITTORALIS. Strapwort. Plate 12 
Corrigiola litoralis L. Sp. PI. 271 (1753)!; Withering Bot. Arr. ed. 2, 322 (1787); Smith Eng. Bot. 
no. 668 (1799); FI- Frit. 339 (1800)!; Eng. Ft. ii, 113 (1824); Syme Eng. Bot. vii, 177 (1867); Rouy Ft. France 
xii, 11 (1910). 
leones: — Smith Eng. Bot. t. 668; Sibthorp and Smith FI. Graec. t. 292. 
Camb. Brit. FI. iii. Plate 12. (a) Flowering shoot. ( b ) Young shoot, (c) Ground-leaves. ( d ) Portion 
of stem and leaves (enlarged). ( e ) Flowers (enlarged). (/) Fruit (enlarged). Devonshire (F. J. H.). 
Exsiccata : — Billot, 19; Dickson, xiv, 10; Fries, xii, 59; Reichenbach, 482; Schultz, 443 bis ; Thielens 
et Devos, i, 7 ; Todaro, 430. 
Annual. Shoot rather glaucous. Branches prostrate or decumbent, slender, up to 45 cm. long. 
Stipules acuminate, membranous. Laminae linear to narrowly oboval. 
Pedicels short. Flowers July to September. Calyx small, with 5 segments, 
segments with scarious margins, persistent. Petals white, ligulate, a little 
longer than the sepals. Stigmas 3, sessile, very small. Capsule oval, 
brown, with 3 longitudinal lines. 
Sandy and shingly places near the sea, subject to inundation of 
fresh water, in Cornwall and Devonshire ; locally abundant. 
Denmark, Germany, Holland, Belgium, France, central Europe, Russia, 
southern Europe ; Africa ; Asia Minor ; central America (introduced). 
Family 2. DIANTHACEAE 
Dianthaceae Tanfani in Parlatore’s Ft. Ital. ix, 239 (1892); Caryophylleae Jussieu Gen. PI. 299 (1789) 
emend.; Bentham and Hooker Gen. PI. 141 (1862); Rouy et Foucaud Ft. France iii, 86 (1896); Caryophyllaceae 
Reichenbach Handb. 296 (1837) emend.; Robinson and Fernald in Gray’s New Manual 377 (1908). 
Perennial or annual herbs, rarely undershrubs. Stem more or less thickened at the nodes. 
Leaves usually exstipulate (stipulate in Polycarpon, Spergula , and Spergularia), petioled or sessile, 
opposite and decussate or rarely in fours, often more or less connate at the base, usually entire. In- 
florescence cymose, often a dichasial cyme. Flowers usually heterochlamydeous, rarely monochlamydeous 
by reduction, usually monoclinous, cyclic, usually obdiplostemonous, rarely partly or almost entirely 
dioecious, usually protandrous. Sepals n , persistent. Petals n, inserted on the disc, rarely absent. 
Disc of hypogynous or hemi-perigynous glands present and usually nectiferous. Stamens inserted 
on the disc, usually n + n, outer whorl usually antipetalous and the first to dehisce, rarely the inner 
whorl (and still more rarely some of the outer whorl also) suppressed ; filaments usually free ; 
anthers dehiscing longitudinally. Ovary superior, syncarpous, of n or n—i or n — 2 carpels, carpels 
usually antipetalous ; carpellary septa present and united to a central column when young, usually 
breaking down before maturity and leaving the ovules on their detached margins, occasionally (as 
in the higher or more specialised forms) more or less persistent in the mature fruit. Gynophore 
often (in the specialised forms) more or less distinct. Styles usually absent or free. Stigmas as 
many as the carpels. Placentation usually free-marginal 1 , as described above (but cf. Sileneae). 
Fruit a capsule, unilocular or (especially in Silene) partially septate, dehiscing by as many or by 
twice as many valves or teeth as there are stigmas, rarely (as in Cucubalus baccifer ) succulent. 
Seeds 4 or more to each carpel, rarely (as in Polycarpon ) fewer. Embryo usually curved round the 
endosperm, rarely straight (as in the specialised Diant heae). Cotyledons narrow, (n =5 or 4.) 
1 This term more clearly expresses the facts of development than the usually accepted term “free-central." 
Map 6. C. littoralis occurs 
in Cornwall and Devon- 
shire 
